1899] THE COMPOUND OOSPHERE OF ALBUGO BLITI 233 
ways far more favorable for study ; its nuclei are larger, its peri- 
plasm more abundant, its developmental stages more ‘strongly 
marked, and its antheridial tube larger. Albugo candidus, how- 
ever, has a remarkable coenocentrum, which will be much easier 
ostudy than that of A. Bliti, owing to its much greater size and 
more pronounced stain reaction. A problem of great impor- 
tance lies in a comparative study of the ccenocentra of the 
genus, 
The characteristic massing of the cytoplasm to form a rudi- 
mentary oosphere in 4. Bhiti, differing thus from the vacuolate 
oosphere of A. candidus, is not a wider divergence than might be 
expected in different Species; nor is the variation in the cceno- 
‘eatrum more than what might be regarded as a specific differ- 
ence. If such variations are found to be more marked in other 
Species the Way may be clear to trace the relationship between 
plants with one oosphere and those with several oospheres in 
tach ogonium ; between forms which differentiate their peri- 
F plasm alter the manner of Vaucheria, and others that follow the 
tof the Saprolegniacez in a parietal rather than a central 
massing. It must be left to future research to make clear the 
ae that must exist between the multiple fertilization 
| ed by Albugo Bliti and simple acts of fusion between 
; “xual elements, 
Ttmay be that cytological investigation will show remarkable 
_ Vafiations in 
dei: many respects in this genus, and establish a chain 
wwe forms. The Saprolegniacez are said to range from 
omplete sexual fertilization. Should Albugo 
ly rich in habits the present knowledge of 
much increased. 
METHODs, MATERIALS, AND STAINING REACTIONS. 
© materj 
- al 
Sisteq a 
Pon which this investigation was based con- 
ems, and flower clusters of Amaranthus retro- 
, ba. oA. hybridus L. bearing the fungus. It was collected 
SES the my = Y,, Columbus, Ohio, and Chicago, III. In all 
Peles seemed to be unquestionably Adbugo Bliti Biv. 
