348 BOTANICAL GAZETTE | NOVEMBER 
stages of synapsis are marked by the same striking peculiarities 
that were met with in Convallaria (fig. 30). In just the same 
way large globules of some deeply stainable matter accumulate 
on the outside of the contracted mass, appearing as if expelled 
from it. They seem, however, to be distinct from the nucleolus, 
as there is no apparent fragmentation nor budding of the latter. 
The nucleolus throughout the whole process remains of exactly 
the same size and regular contour, and with the little wart-like 
attachment undisturbed. 
The spirem stage is much shorter than in Convallaria. The 
contracted linin network gradually begins to expand until the 
threads are again spread out beneath the membrane. Here 
again we notice a decided change in the structure of the linin 
thread, just as was the case in Convallaria. It is no longer s0 
slender, and provided with such large granules, nor is it so con- 
spicuously in the form of a network. The spirem is composed 
of a few rather thick linin threads extending in various direc- 
tions around the nucleolus, and crossing each other occasionally. 
In them appear small chromatin granules, which, however, af 
much smaller and more regular in size than those present before 
synapsis (fig. 37). The whole process, therefore, is exactly 
comparable with that in Convallaria. 
FIRST NUCLEAR DIVISION OF THE MOTHER-CELL. 
The stages in the nuclear development preparatory to the 
first nuclear division are not nearly so marked as in Convallana. 
The first indication of division is found in the gradual massing 
together of the chromatin into a number of irregular masses 
simulating those found just before synapsis. The difference a 
in their larger and more equal size. Just at the time when ef 
nuclear membrane is disappearing the number of these gee 
of chromatin may be determined approximately. Proba if 
fourteen or sixteen is the correct number. They S007 ae 
lie together in pairs, in which case two may be easily mist : 
for one (fig. 32). During the later stages the two pan 
each pair seem to lose their identity, so that when on the sp! 
