THE DEVELOPMENT AND ANATOMY OF SARRACENIA 
PURPUREA.* oe 
CONTRIBUTIONS FROM THE BOTANICAL LABORATORY OF THE 
JOHNS HOPKINS UNIVERSITY. No. 5. 
FORREST SHREVE. 
(WITH PLATES III-V) 
THE work of which the results are here given was undertaken at 
the suggestion of Dr. D. S. JoHnson, and has been carried out at the 
Biological Laboratory of the Johns Hopkins University. I wish 
here to express my thanks to Dr. JoHNson for much advice and 
helpful criticism in connection with this work, and to express to 
Professor Witu1am K. Brooks my appreciation of his interest and 
encouragement, I also wish to thank my fellow-student Mr. SAMUEL 
RITTENHOUSE for his kindness in gathering material for me during 
my absence from Baltimore. 
The material worked upon was obtained mainly at Glenburnie, 
Maryland, near Baltimore. Most of it was fixed in the field; and 
of several killing reagents tried 1 per cent. chrom-acetic and Carnoy’s 
mixture were the most satisfactory. Preparations mere made by 
ordinary paraffin method and stained with the Flemming triple stain 
or with cyanin and erythrosin. 
DEVELOPMENT OF THE FLOWER. 
The earliest stage observed in the development of the flower was 
in material gathered August 30. There are then to be seen the 
primordia of the three bracts, the five sepals, and the five petals, 
which have apparently arisen in the order named. Lying just 
within the edges of the petals are the staminal primordia, as yet mere 
papillae, and within them is a flat surface with slight elevation at 
the center. A somewhat later stage than the last shows progress in 
the development of the stamens, which now appear as ten groups of 
protuberances lying in the position before noted (fig. 2). Each 
« Dissertation submitted to the Board of University Studies of the Johns Hopkins 
University for the degree of Doctor of Philosophy. 
a] . [Botanical Gazette, vol. 42 
