II0 BOTANICAL GAZETTE [avoust 
main placental outgrowth sends out two lateral outgrowths so as to 
resemble in cross section a letter T, in which the arms have been 
bent downward. Each pair of adjoining lateral outgrowths 
closely appressed and directed backward toward the angles of the 
ovary. In the lower part of the ovary the adjacent lateral out- 
growths fuse, but do not extend to the bottom, and in the upper part 
they do not reach the wall of the ovary as do the main placental — 
outgrowths. Upon the edges of the lateral outgrowths and upon — 
the surfaces lying next the main outgrowths are borne the ovules 
(fig. 15). The ovules at the base and top of the ovary lie ea 
the intermediate ones having intermediate positions according : 
their place in the ovary. 
The summits of the carpellary leaves broaden and coalesce, and 
grow out in a direction radial to the axis of the flower, so that while 
their basal parts form the capsule and the stalk of the style, the tips 
form the umbrella of the style (fig. 3). The tip of each carpellary 
leaf organizes a very definite growing-point (fig. 28), and the portion 
between the tips nearly keeps pace in growth. Upon the ventral 
surface of each tip, just before it completes its growth, is formed the 
protuberance which bears the stigmatic surface. F 
The appearance of the primordia of the ovules upon the placentae — 
takes place from the point opposite the angle of the ovary wall, 
where the adajcent lateral outgrowths meet, successively toward — 
the angle formed by the lateral outgrowth and the main outgrowth 
(fig. 19). In vertical direction the development proceeds from the — 
epidermal cells and the accompanying anticlinal division of the 
epidermal cells, as is commonly the case. When the ovule first — 
protrudes from the placenta there is no suggestion of a sporogenous 
cell. At this stage of development the winter rest intervenes. The 
first suggestion of a sporogenous cell comes with the enlargement of 
a single subepidermal cell, which is the megaspore mother cell (fig. 1)- : 
In three cases out of many hundreds examined there were two mother — 
cells lying side by side. There is no tapetal cell. The bending by 
which the ovule becomes anatropous begins at once, and is quite 
