122 BOTANICAL GAZETTE [Aucusr 
The axillary buds of Sarracenia are commonly very small and 
consist of growing-point and the primordium of a single leaf, com- 
pletely covered and protected by the sheathing leaf base. An 
occasional axillary bud develops, the first two leaves being opposite 
and on the opposite sides of a line connecting the growing-point 
of the bud with the center of the shoot. There is thus brought 
about a branching of the rhizome, which by frequent repetition gives 
rise to large clusters of individuals. 
The anatomy of the rhizome of S. purpurea has been described 
by ZIPPERER in sufficient detail, since it presents no unusual fatures. 
He has also given a correct account of the growing-point of the root 
and the development of the vascular tissue of the root: the growing- 
point being of the type in which the cap and three tissue layers are 
all derived from a common group of initials; the early order of 
vascular bundles being triarch. 
Root hairs are few upon the roots of plants growing in a saturated 
substratum in the open, but are abundant in seedlings grown in 
highly saturated sphagnum, and in adult plants in the open which 
are growing in a substratum merely moist. Mycorhiza has not been 
observed in S. purpurea in the vicinity of Baltimore, although fungal 
threads have been found covering the root of seedlings grown under 
the conditions previously mentioned and penetrating the epidermis. 
MacDoveat’ (1899) has described penetration of the epidermis 
by hyphae in adult plants without committing himself as to their 
mycorhizal nature. 
SUMMARY. 
1. The flowers of Sarracenia purpurea are axillary, perfect, 
hypogynous, and radially symmetrical. The stamens are seventy — 
to eighty in number and arise in ten groups. There are four micro- — 
sporangia. There is a double layer of binucleate tapetal cells, 
derived from the primary archesporium. There are three to five 
parietal layers. The tetrad division is simultaneous; the micro- 
spore nucleus divides before the dehiscence of the anthers. The 
reduced number of chromosomes is twelve. 
2. In the ovule there is a single archesporial cell, which is the 
megaspore mother cell. There is no tapetal cell. The ovule is 
7 Symbiotic saprophytism. Annals of Botany 13:1-47. 1899. 
