1906] COOK—CUBAN ._NYMPHAEACEAE 387 
pubescens, Brasenia purpurea, and Cabomba piauhiensis all have the 
same general « character as Nymphaea advena, except that they pos- 
sess suspensors. ‘They also show some resemblance (which may be 
superficial) to the embryo of Sparganium simplex as described by 
CAMPBELL (3). They also resemble the embryos of Naias flexilis 
and Zannichella palustris as described by CAMPBELL (1), Potamoge- 
ton foliosus and P. natans as described by WIEGAND (27) and Hot- 
FERTY (10), and Limnocharis emarginata as described by HALL (9g) 
except that they do not possess the large basal cell of the suspensor, 
The late development of the suspensor in Nymphaea advena ( ?) of 
Cuba and N. advena of the north, as previously described by me, 
may confirm in some measure COULTER and CHAMBERLAIN’S view 
that the Alisma-type of monocotyledonous embryo is primitive and 
that the suspensor in the Pistia type has been suppressed. 
It will also be noted that the origin of the cotyledonary lobes in 
all cases is from a crescent-shaped cotyledonary ridge about the 
larger part of the embryo, and that my results coincide with the con- 
clusions reached by Lyon, ScHAFFNER, and York. This point in 
Conarn’s studies is illustrated by a single figure (48, d) of Castalia 
caerulea. This figure agrees exactly with fig. 30 and text jig. 2, d, e 
of my first paper, with Lyon’s fig. 10, and Yorx’s fig. 33 of Nelumbo. 
However, my figures are parts of series which illustrate the true 
~ Monocotyledonous character of the embryo; and this monocotyle- 
donous character of C. odorata was afterwards demonstrated by 
SCHAFFNER who dissected the embryos out of their sacs. 
RICHARD (21) referred to the embryos of the grasses, Nelumbo, 
Ruppia, Hydrocharis, and Zostera as embryons macro podes, and 
SCHAFFNER (26) confirms this view and expresses the opinion that the 
Massive expansion and lobes of Halophila, Ruppia, Zostera, Nelumbo, 
Nymphaea, and Castalia are not homologous with the cotyledons, 
but are specially developed absorbing organs. 
The figures of this paper show a striking resemblance to the 
figures of anomalous dicotyledons by Lewis (16) and Mortrier 
(20). However, the flowers show more monocotyledonous than 
dicotyledonous characters, while only the leaves may be considered 
4S uniformly presenting dicotyledonous characters. It appears, 
therefore, that the Nymphaeaceae can be more properly classed as 
