240, ON THE GLOSSOPHAGINE. 
The origin of the Glossophagine is easily traceable to the group denominated by 
Peters the Vampyri. But the division between the genera composing the Vampyri is 
of a character to suggest two groupings at least, and the term Vampyri is best used in a 
restricted sense. Indeed, it is a small cluster of four genera only ( Vampyrus, Macrotus, 
Schizostoma and the aberrant Hemiderma), which possess a large, triangular, first upper 
premolar and an inflated, weak periotic region. . 
Of the second group (Phyllostomi), of which Phyllostoma is the type, I have imper- 
fect knowledge—haying studied besides this form the genera Lonchorhina and Lophostoma. 
But they agree in having the first upper premolar small and acicular, a peculiarity I find 
figured in Geryais (Hxp. du Sud.) as characteristic of Tylostoma and Monophyllum 
(Dolichophyllum). 1 infer that Trachyops, Phylloderma and Mimon are members of this 
group from Dobson’s statement (Br. Cat. Chir.) that they resemble Phyllostoma.. I have 
no satisfactory knowledge of the periotic region in this group, but can say that it is boldly 
defined, concave, and not inflated in Phyllostoma, Lonchorhina and Lophostoma. 
Now it has been seen that the Glossophagine yield two groups—that of the Glosso- 
phagi and that of the Lonchoglossi. In my judgment these do not haye a common origin. 
The Glossophagi agree with the Vampyri as above restricted in the shape of the first 
upper premolar and the inflated periotic region, while the Lonchoglossi are much nearer 
the Phyllostomi. Chenycteris possesses a triangular premolar (with large denticles) 
and a moderately truncate concaye periotic region, but its other characters, taken as a 
whole, connect the form intimately with the Glossophagi. 
The taxonomic value of the terminal cartilage can be determined only by the 
examination of extended series. At first I had inferred that the shapes of the cartilages 
of the fourth and fifth digits were of considerable value. But inspection of the largest 
number of individuals of the most common species—namely, Glossophaga soricina—gave 
me an impression that they were really variable structures ; thus in one individual from 
Costa Rica they were both spatulate ; in another from Bahama Islands they were both 
aciculate ; and yet in a third specimen from the last-named locality the fourth digit was 
spatulate and the fifth aciculate. Nevertheless the variability itself is of interest and I 
have, therefore, figured the cartilages, believing that after extended observation they may 
assist In more firmly defining the minor groups of species than is now the case. 
GLOSSOPHAGA. 
Upper incisors in a continuous row. Length of forearm not exceeding 56 mm.; thumb, 
8 mm.; calear present; the tail is short with free tip on the dorsum of the interfemoral 
membrane. Proencephalon creates an eminence on brain case ; fronto-maxillary inflation 
conspicuous ; mastoid process small. 
Dental formula: 1. +— ¢. +— prm. 4—m., $= 21. 
