NOTES ON THE CANID# OF THE WHITE RIVER OLIGOCENE. BAT 
The distal end is large and has a well-developed trochlea, which is much more strongly 
convex than in Canis and of a different shape, the modern genus haying here a trochlea 
which is more like that of a phalanx than of a typical metacarpal. In Daphwnus, but 
not in Canis, there is a well-defined palmar carina, and the lateral processes for ligamen- 
tous attachment are more prominent than in the recent type. 
The second metacarpal is much longer and stouter than the first, though very short 
with reference to the size of the animal and to the length of the other segments of the 
fore limb. The proximal end is not much expanded transversely, but has a great dorso- 
palmar extension, the head projecting much farther behind the plane of the shaft than in 
Canis. The facet for the trapezoid is less concave transversely than in the modern genus 
and is of more uniform width, narrowing less toward the palmar side ; the ulnar border 
rises more above the head of me. iii and has a more extensive contact with the magnum. 
Though larger than in the recent Canide, this contact with the magnum is much smaller 
than in existing felines, and is of about the same proportions as in the early sabre-tooth, 
Hoplophoneus. The combined facets for the magnum and for me. iii form a broad, 
curved band upon the ulnar side of the head, which is made slightly concave to receive 
the adjoining metacarpal. No distinctly marked facet for the trapezium is visible upon the 
radial side. The shaft is short, weak, of transversely oval section, and is arched toward 
the dorsal side. The distal end is expanded, and made broad by the large, rugose pro- 
cesses for the attachment of the lateral metacarpo-phalangeal ligaments, processes which 
are much better developed than in Canis. The distal trochlea is of a quite different shape 
from that seen in the modern genus, being narrower, higher and of more nearly spherical 
outline, and is demarcated from the shaft by a deep depression, such as does not occur in 
the existing members of the Canide. The palmar carina is prominent and thins to a 
narrow edge. 
The third metacarpal is incomplete in the only manus found in the collection 
(D. felinus, No. 11425, Pl. XX, Fig. 17) as it lacks the distal end. The portion pre- 
served is, however, as long as the whole of me. ii and the complete bone was evidently 
considerably longer. The shape of the proximal end is much as in Canis, except for the 
relatively greater dorso-palmar diameter. The magnum facet is narrow, but deep, some- 
what concave transversely and strongly convex antero-posteriorly, but less so than in 
existing dogs. The facet on the radial side for me. ii is larger, more oblique and more 
prominent, and is more extensively overlapped by me. ii than in the latter, and the 
surface for me. iv, while not so deeply concave, is larger. When the third and fourth 
metacarpals are placed together in their natural positions, it is seen that the former rises 
higher proximally than the latter and has a contact with the radial side of the unciform, 
which, though narrow, is larger thanin Canis. The shaft is somewhat more slender than 
