384 NOTES ON THE CANID# OF THE WHITE RIVER OLIGOCENE. 
VI. Tse Manus (Pl. XX, Fig. 23). 
By a fortunate discovery of Mr. Hatcher’s, 1am enabled to give an account of an 
almost complete carpus belonging to Cynodictis, which has hitherto been entirely 
unknown. 
A scapho-lunar is present, formed by the coalescence of the scaphoid, lunar and 
central, which distinguishes Cynodictis from the creodonts. This bone resembles that of 
Canis in general character, but displays quite a number of differences in points of detail, 
and these differences are, at the same time, approximations to the structure found in 
Daphenus. The seapho-lunar has a very small vertical (proximo-distal) diameter, 
especially on the radial side, where it thins away to a mere edge, the facets for the radius 
and the trapezium almost meeting. As compared with the corresponding carpal of Canis, 
this bone has a somewhat greater transverse and smaller dorgo-palmar diameter. The 
radial facet is simply convex both transversely and antero-posteriorly, and has not the 
saddle-shaped extension at the interno-palmar angle which is found in the recent dogs. 
This facet descends quite low upon the dorsal side of the bone, as is also the case in 
the modern plantigrade and semiplantigrade carnivores. The hook-lke process which 
arises from the postero-internal angle of the scapho-lunar is much shorter and less mas- 
sive in every dimension than that of Canis. Another difference from the modern genus 
consists in the absence of any distinct articular surface for the pyramidal, the facet for 
the radius and that for the unciform almost coming into contact along the ulnar side of 
the bone. 
On the distal side of the scapho-lunar are four facets, for all the carpal elements of 
the distal row. That for the unciform is relatively smaller than in Canis, and is con- 
fined to a narrow strip near the ulnar border; the magnum facet is much the same as in 
the modern genus, but is somewhat more oblique in position. The surface for the tra- 
pezoid is fairly large and keeps more nearly parallel with that for the magnum than in 
the recent dogs, while the trapezium facet is small and of almost circular shape. 
The pyramidal is a very different-looking bone from that of the modern dogs, 
being broad, depressed and scale-like in shape; its vertical (or proximo-distal) diameter is 
very small and relatively much less than in Canis, and there is no such process from 
the ulnar side of the bone as in the latter, in which the pyramidal articulates with the 
head of the fifth metacarpal by a much more extensive facet than in Cynodictis. 
The recent viverrines have the pyramidal shaped very much as in the White River 
genus. The proximal surface is divided into two narrow and somewhat concave facets 
for the ulna and pisiform respectively, of which the latter is slightly the larger. On 
the distal side is a single large and concave facet for the unciform, and posterior to this 
