96 NOTES ON THE CANIDEH OF THE WHITE RIVER OLIGOCENE. 
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navicular is transverse, and owing to the elongation of the neck of the astragalus, it is 
carried so far distally that it can have no contact with the caleaneum, the astragalus 
articulating with the cuboid. The astragalar surface is concave, but somewhat less so 
than in Canis, and the facet for the cuboid is small and confined to the dorsal moiety of 
the fibular side. The distal end displays the usual facets for the three cuneiforms, which 
do not require any particular description. 
The entocuneiform has much the same shape as in Canis, elongate in the proximo- 
distal diameter, but very narrow and much compressed. The nayicular facet is rela- 
tively smaller than in the modern genus and there is no such distinct facet for the meso- 
cuneiform. The distal surface, for the head of the first metatarsal, is no wider but much 
more deeply concave than in Canis. 
The mesocuneiform is a minute bone and, as in the fissipede Carnivora generally, its 
vertical or proximo-distal diameter is much less than that of the adjoining ento- and 
ectocuneiforms, forming a depression or recess in the distal row of the tarsus, into which 
the head of the second metatarsal is tightly wedged. The only articular surfaces visible 
on the mesocuneiform are the proximal and distal, for the nayicular and the second meta- 
tarsal respectively. 
The ectocuneiform is much the largest of the three. Compared with that of Canis, 
it is narrower in proportion to its height and is also less extended in the dorso-plantar 
dimension, but the projecting process from the plantar surface is even more prominent, 
and is more thickened and club-shaped at the free end. On the tibial side is a minute 
facet (not double as in Canis) for the side of mt. ii. The facet for the cuboid is much 
smaller than in the modern dogs and is confined to the dorsal border, while at the infero- 
external angle of the bone is a minute facet for the head of mt. iv, which is not repre- 
sented in Canis. The distal end of the ectocuneiform is taken up by a facet for mt. 111, 
which is less concave and has a shorter plantar prolongation than in the modern genus. 
The metatarsus consists of five well-developed members. Unfortunately, there is 
not a single complete metatarsal preserved in connection with any of the specimens, but 
enough remains to show that these bones were much longer and stouter than the meta- 
carpals, and that the disproportion in size and length between ‘the fore and hind feet 
was much greater than in the recent dogs and quite as great as in many viyerrines, such 
as Herpestes and Paradoxurus or as in Daphenus. 
The first metatarsal is sufficiently weli preserved to indicate that the hallux was 
well developed and functional, though somewhat more reduced than in Daphenus, or in 
such recent viverrines as Cynogale or Paradoxurus. The head bears a narrow, convex 
facet for the entocuneiform and upon its tibial side is a large, rugose prominence for the 
attachment of the lateral ligament. The shaft is very slender and is arched slightly 
