Gleicheniaceae; Hymenophyllaceae 33 



brown jointed hairs while young; stipes pale brown, terete; jointed, brown, soon deciduous, pinnately veined, the veins 



fronds sometimes more than 1 m. long, firm; ultimate 2-A times forked, slighdy raised on both sides; sori small, 



branches (pinnae) chartaceous, linear- to oblong-lanceolate, rounded, solitary near the middle portion of the uppermost 



pinnatiparted, 15-30 cm. long, 3-7 cm. wide, gradually acute, vein. Oct.-Jan. Dry thin woods and thickets; Honshu 



obtuse at the tip, sessile, green above, glaucous beneath; seg- (Iwaki and Echigo Prov. westw.), Shikoku, Kyushu; common. 



ments many, nearly horizontally spreading, linear, 3^ mm. Korea, Ryukyus, Formosa, China, and widely distributed 



wide, obtuse to retuse, entire, slightly hairy beneath, the hairs in die Tropics and subtropics, also in Australia. 



2. GLEICHENIA J. E. Smith Urajiro Zoku 



Rhizomes long-creeping, firm, scaly while young; fronds large, stipitate, pinnate or falsely forked; pinnae bipinnatiparted; 



veins once forked; sori superficial, with about 4 sporangia. About 10 species, Malaysia to India and Japan, 1 in tropical 



America. 



lA. Scales ciliate; pinnules nearly horizontally spreading, glaucous and glabrous beneath; ultimate segments narrowly oblong, obtuse; 



stipes terete 1- l^- sfatica 



IB. Scales entire; pinnules obliquely spreading, slightly glaucous with scattered glandular hairs beneath; ultimate segments broadly linear, 



cuspidate; stipes flattened and with 2 longitudinal ridges to the base 2. G. laevtssima 



1. Gleichenia japonica Spreng. Polypodium glaucum the anterior branches of the veins. Nov .-Jan. Dry, thin 



Houtt.; G. glauca (Houtt.) Hook., non Sw.; Dicranopteris woods; Honshu (Iwaki and Echigo Prov. westw.), Shikoku, 



glauca (Houtt.) Underw.; Diplopterygium glaucum (Houtt.) Kyushu; common. Korea, Ryukyus, Formosa, China, the 



Nakai; Hicriopteris glauca (Houtt.) St. John Urajiro. Tropics of Asia, Australia, and Polynesia. 



Rhizomes long-creeping, terete, 5-6 mm. across, wiry, densely 2. Gleichenia laevissima Christ. G.k.iusiana Makino; 



scaly while young; stipes terete; scales lanceolate-deltoid, 6-7 Dicranopteris laevissima (Christ) Nakai; Diplopterygium lae- 



mm. long, caudate, ciliate, lustrous, dark brown; fronds scaly vissimum (Christ) Nakai; Hicriopteris laevissima (Christ) 



while very young, die dormant terminal bud densely scaly, St. John Kaneko-shida. Closely allied to the preceding; 



rufous to dark brown, ciliate; pinnae chartaceous, opposite, scales brown to yellow-brown, membranous, entire, glabrous; 



50-100 cm. long, 20-30 cm. wide, bipinnatiparted, green, stipes widi a narrow flat face and 2 longitudinal ridges to 



slighdy lustrous above, glaucous beneath, glabrous; pinnules the base; pinnules obliquely spreading, the ultimate segments 



many, horizontally spreading, linear-lanceolate to broadly broadly linear, acute or cuspidate, green above, slightly glau- 



linear, 15-30 mm. wide, sessile, pinnatiparted, the ultimate seg- cous and with short subclavate minute glandular hairs be- 



ments spreading, narrowly oblong, obtuse; costa impressed neath. Oct.-Nov. Kyushu; rare. China, Indochina, and 



above, the costules slender; veins once forked; sori median; on Philippines. 



Fam. 11. HYMENOPHYLLACEAE Koke-shinobu Ka Filmyfern Family 



Epiphytic or terrestrial ferns; rhizomes slender, long-creeping, nearly naked or with jointed hairs; fronds usually small, thin, 

 without stomata, usually with circinnate vernation, very rarely straight in extremely reduced fronds; veins nearly always free; 

 sporangia in marginal sori elevated with a slender receptacle surrounded by a cup-shaped sometimes 2-valved indusium; an- 



nulus complete; spores tetrahedral to globose. About 34 genera, with about 650 species, chiefly in the Tropics and subtropics, 



few in colder regions. 



lA. Indusium 2-valved; rhizomes very scantily hairy; fronds fulvous to brownish green. 



2A. Ultimate segments of the blades entire 1. Mecodium 



2B. Ultimate segments of the blades toothed 2. Hymenophyllum 



IB. Indusium cup-shaped to tubular, entire or shallowly 2-Iobed at mouth; rhizomes prominently hairy; fronds green. 

 3A. Rhizomes long-creeping, remotely leafy. 

 4A. False veinlets absent. 



5A. Fronds pinnately compound 3. Vandenboschia 



5B. Fronds palmately lobed to parted 4. Gonocormus 



4B. False veinlets present 5. Crepidomanes 



SB. Rhizomes short, erect to ascending, densely leafy 6. Sclenodesmium 



1. MECODIUM PresI Koke-shinobu Zoku 



Epiphytic or growing among mosses on rocks; rhizomes very slender, long-creeping, thinly hairy to subglabrate; fronds 

 usually thin, pinnately compound, the ultimate segments entire, 1-veined; indusium 2-valved, usually nearly to the base, the re- 

 ceptacle included. A pan tropic genus comprising about 100 species. 



lA. Stipes prominently winged. 



2A. Plants usually 10-30 cm. high; ultimate segments of frond 1-2 mm. wide; wings of stipes entire to slightly crispatc-undulate. 



3A. Ultimate segments of blades 1-1.5 ram. wide; main rachis 1.5-2 mm. wide inclusive of the wings; fronds narrowly to broadly 



lanceolate 1. M. flexile 



SB. Ultimate segments of blades about 2 mm. wide; main rachis 2-3 mm. wide inclusive of the wings; fronds oblong to ovate-oblong. 



2. M. badium 

 2B. Plants 5-10 cm. high; ultimate segments of frond 0.5-1 mm. wide; wings of stipes strongly crispatc-undulate 3. M. riul^iuense 



