34 



Hymenophyllaceae 



IB. Stipes not or scarcely winged. 

 4A. Stipes and nerves on underside of blades and rachis with yellow-brown flexuous hairs; plants about 5 cm. high. . . 4. M. oligosorum 

 4B. Stipes and blades glabrous or nearly so; plants taller. 



5A. Pinnae disposed at an acute angle on the rachis, simple or the segments 2-5, elongate, 1.2-1.7 mm. wide 5. M. wrightii 



53. Pinnae disposed at an obtuse angle on the rachis, the segments narrower, 0.7-1 mm. wide. 



6A. Fronds 10-25 cm. long; stipes narrowly winged 6. M. polyanthos 



6B. Fronds 2—5 cm. long; stipes not winged 7. M. paniculiflorum 



1. Mecodium flexile (Makino) Copel. Hymenophyl- 



lum flexile Makino O-koke-shinobu, Nachi-koke-shinobu. 



Rhizomes long-creeping, filiform, remotely leafy; fronds 10- 

 30 cm. long; stipes winged except at base; blades narrowly 

 to broadly lanceolate, 7-20 cm. long, 1.5-3 (-4) cm. wide, usu- 

 ally narrowed at tip, 2- or 3-pinnatiparted, the rachis 1.5-2 

 mm. wide inclusive of the entire or slighdy crispate wing; pin- 

 nae rather numerous, narrowly ovate, somewhat oblique, 

 pinnately to bipinnately parted, the pinnules 1-1.5 mm. wide; 

 sori few on each pinna, the valves orbicular to slightly de- 

 pressed, entire, 1.5-2 mm. across. Among mosses; Honshu 



(Izu Prov. through Tokaido to Kinki Distr.), Shikoku, Kyu- 

 shu. Formosa. 



2. Mecodium badium (Hook. & Grev.) Copel. Hytne- 

 nophyllutn badium Hook. & Grev.; H. crispatum sensu auct. 

 Japon., non Wall. Oni-koke-shinobu. Plant very spar- 

 ingly pilose, mosdy while young; rhizomes long-creeping, 0.5- 

 0.7 mm. across; fronds oblong to ovate-oblong, 15-30 cm. 

 long, 4-7 cm. wide; stipes 5-10 cm. long, 2-3 mm. wide in- 

 clusive of the entire sometimes slightly crispate wings; blades 

 ovate to narrowly so, usually narrowed at tip, 3-pinnatiparted, 

 the rachis winged, 2-3 mm. wide; pinnae obliquely spreading, 

 usually somewhat elongate; ultimate pinnules or segments en- 

 tire, broadly linear, about 2 mm. wide, obtuse; indusia de- 

 pressed-orbicular, the valves obsoletely toothed on upper mar- 

 gin or entire, 2-3 mm. wide. Among mosses on rocks and 



on tree trunks; Honshu (Kii Prov.), Shikoku, Kyushu; rare. 

 Formosa, China to Malaysia, and India. 



3. Mecodium riukiuense (H. Chr.) Copel. Hymeno- 



phyllum nu\iuense H. Chr. Ryukyu-koke-shinobu, Oki- 



NAWA-KOKE-SHiNOBU. Rhizomcs Creeping, nearly filiform; 



fronds 5-10 cm. long, 17-30 mm. wide; stipes at first with an 

 entire, soon crispate-undulate wing on both sides; blades nar- 

 rowly ovate, sometimes somewhat elongate and narrowed at 

 tip, 3-pinnanparted, the rachis winged; ultimate pinnules or 

 segments narrow, linear, 0.5-1 mm. wide, obtuse; valves of 

 indusia ovate to elliptic, few-toothed on upper margin, about 



1 mm. wide. Shikoku, Kyushu (s. distr.); rare. Ryu- 



kyus. 



4. Mecodium oligosorum (Makino) H. Ito. Hymeno- 



phyllum oligosorum Makino Kiyosumi-koke-shinobu. 



Rhizomes, rachis, and costa of the fronds rufous-pubescent; 

 rhizomes capillary, long-creeping, sparsely branched; fronds 

 small, about 5 cm. long; stipes filiform, 1-3 cm. long, not 

 winged; blades broadly to narrowly ovate, 2-4 cm. long, 

 about 1.5 cm. wide, obtuse, bipinnately parted, the rachis 



winged; pinnae obliquely spreading, palmately to pinnately 

 parted, the ultimate segments obtuse, entire; indusia orbicular, 

 about 1.5 mm. wide, the valves obsoletely few-toothed on up- 

 per margin. Among mosses on rocks in damp woods; Hon- 

 shu (Kanto Distr. and Mino Prov. westw.), Shikoku, Kyu- 

 shu. 



5. Mecodium wrightii (v. d. Bosch) Copel. Hymeno- 



phyllum wrightii v. d. Bosch Koke-shinobu. Delicate, 



very thinly hairy, especially while young; rhizomes capillary, 

 branched, forming a loose mat; fronds small; stipes 7-20 mm. 

 long, glabrous, narrowly winged in upper part; blades deltoid- 

 ovate to narrowly so, 1.5-3 cm. long, 1-1.5 cm. wide, bipinnati- 

 parted, the rachis winged; pinnae rather few, ascending, di- 

 vided or simple in the upper ones; ultimate segments more or 

 less elongate, 1.2-1.7 mm. wide, entire, retuse or sometimes 

 rounded at apex; indusia orbicular to broadly ovate, the valves 

 1.2-1.5 mm. wide, entire or obsoletely bilobed on upper margin. 



Damp woods in mountains; Hokkaido, Honshu, Shikoku, 



Kyushu. Sakhalin and Korea. 



6. Mecodium polyanthos (Sw.) Copel. Trichomanes 

 polyanthos Sw.; Hymenophyllum polyanthos Sw.; H. blume- 

 anum Spreng.; H. integrum v. d. Bosch; H. fujisanense Na- 



kai HOSOBA-KOKE-SHINOBU, HoSOBA-HIME-KOKE-SHINOBU. 



Fronds 10-25 cm. long; rhizomes filiform to capillary, hairy, 

 more especially while young; stipes 2-10 cm. long, narrowly 

 winged, with a tuft of dark brown straight hairs at base; 

 blades broadly lanceolate to ovate, 5-17 cm. long, 2-4 cm. 

 wide, 2- or 3-pinnatiparted, the rachis narrowly winged; pin- 

 nae usually many; pinnules short, entire or few-parted; ulti- 

 mate segments narrow, usually elongate, about 0.7-1 mm. 

 wide, obmse; indusia broadly ovate to suborbicular, about 1 



mm. long, the valves entire, obtuse to rounded at apex. 



Honshu (Iwashiro and westw.), Shikoku, Kyushu; more 



common southward. Ryukyus, Formosa, widely distributed 



in the tropics and subtropics. 



7. Mecodium paniculiflorum (Presl) Copel. Hymeno- 

 phyllum paniculiflorum Presl Hime-koke-shinobu, Foji- 



KOKE-sHiNOBU. Closely allied to the preceding; rhizomes 

 capillary, branched, thinly brown-hairy at the tip; fronds 2-5 

 cm. long, nearly glabrous; stipes capillary, 1-2 cm. long; blades 

 ovate to oblong-ovate, 1.5-3.5 cm. long, 1.2-1.5 cm. wide, ob- 

 tuse, the rachis winged; pinnae ovate to ovate-deltoid, 2- to 

 9-parted, obliquely spreading, the segments emarginate, about 

 1 mm. wide; indusia orbicular, the valves entire, about 1 mm. 



wide. ^Mountains; Honshu (centr. and Kanto Distr.); rare. 



sw. China, Malay Peninsula, and Malaysia. 



2. HYMENOPHYLLUM J. E. Smith Koya-koke-shinobu Zoku 



Epiphytic or terrestrial, with slender creeping scantily hairy rhizomes; fronds small, thin, pinnately compound, the ulti- 

 mate segments toothed; false veinlets absent; indusia 2-valved nearly or to the base; receptacle not or only slighdy exserted. 



About 25 species, of wide distribution, except in cold regions. 



lA. Fronds 3-pinnatiparted, ovate to broadly lanceolate, sparingly brownish hairy on veins and costas on underside and on stipes; ultimate 



segments about 1 .2-1 .7 mm. wide \. H. barbatum 



IB. Fronds 2-pinnatiparted, lanceolate, nearly glabrous; ultimate segments 2-3 mm. wide 2. H. simonsianum 



