38 



Pteridaceae 



2. MICROLEPIA Presl Fumoto-shida Zoku 



Terrestrial, usually hairy, with creeping solenostelic rhizomes; fronds moderate to large, pinnate to decompound, the ulti- 

 mate pinnules obliquely incised; veins free; sori intramarginal, terminating the veinlets; indusia half cup-shaped, glabrous to 

 hispid, the receptacle short; spores tetrahedral. More than 45 species, chiefly in the Tropics and subtropics of the Old World. 



lA. Indusia prominently setose, often ciliate. 



2A. Blades pinnate; rachis uniformly pubescent on both sides I. M. marginata 



2B. Blades bipinnate; rachis glabrous on upper side except upper part 2. M. pseudostrigosa 



IB. Indusia glabrous or with 1 or 2 long hispid hairs near base. 



3A. Blades bipinnate; pinnae 1.5—3 cm. wide 3. M. strigosa 



3B. Blades tripinnate; pinnae 5-10 cm. wide. 



4A. Pinnae lanceolate, the lower about 5 cm. wide; pinnules obtuse to acute 4. M. substrigosa 



4B. Pinnae ovate-deltoid, the lower 5-10 cm. wide; pinnules long-acuminate 5. M. yakusimensis 



1. Microlepia marginata (Houtt.) C. Chr. Polypodium 15-40 cm. wide, abrupdy acuminate, scarcely narrowed at base, 

 marginatum Houtt.; P. marginde Thunb., non L.; Dic\sonia bipinnate, the rachis glabrous and grooved on the stramineous 

 marginalis (Thunb.) Sw.; Davallia marginalis (Thunb.) smooth upperside, densely brownish pilose on the dull purplish 

 Baker; M. marginalis (Thunb.) Bedd. Fumoto-shida. underside; pinnae linear-lanceolate, 1.5-3 cm. wide, long- 

 Fronds evergreen, pilose; rhizomes creeping, 4-7 mm. across, acuminate, thinly hispid on veins especially beneath; pinnules 

 thinly hairy; sripes 30-50 cm. long, more or less scabrous, dull oblong-ovate, 10-20 mm. long, 5-10 mm. wide, acute to obtuse, 

 purple-brown; blades broadly lanceolate, 30-60 cm. long, 15- unequally cuneate at base, sessile, toothed, sometimes lobulate; 

 30 cm. wide, gradually acuminate, pinnate, the rachis uni- sori nearly marginal, in the sinus; indusia membranous, semi- 

 formly pubescent on both sides, dull purpie-brown; pinnae rounded, hyaline, entire, glabrous or sometimes with 1 or 2 



chartaceous-herbaceous, linear-lanceolate, 1.5-3 cm. wide, pin- hispid hairs near base, about 0.7 mm. wide. Honshu (Izu 



natilobed, acuminate, the pinnae of smaller fronds obtuse, sub- Isls., Awa Prov. and westw.), Shikoku, Kyushu. Korea 



sessile, obliquely truncate to broadly cuneate, auriculate or (Quelpaert IsL), Ryukyus, Formosa, China to India, Malaysia, 



with a larger lobe on anterior side at base, spreading setose- and Polynesia. 



hirsute beneath on costas and veinlets, strigose on costas above; 4. Microlepia substrigosa Tagawa. Usuba-ishi-ka- 



ultimate pinnules ovate-deltoid to ovate-oblong, toothed; sori guma. Closely allied to the preceding but the rhizomes 



1-7 on a lobe, submarginal; indusia depressed-orbicular, whit- with dark brown hairs; fronds thinner, the lower pinnae 



ish, prominently setose, often ciliate, about 1.5 mm. wide. broader, lanceolate, about 5 cm. wide; pinnules narrowly ovate 



Honshu (Hitachi and Etchu Prov. westw.), Shikoku, Kyushu; to broadly lanceolate-deltoid, 2-3 cm. long, 6-12 mm. wide, 



common. Ryukyus, Formosa, China to India. obtuse to acute, oblique, short-petiolulate, pinnate to pinnati- 



2. Microlepia pseudostrigosa Makino. M. marginata parted; ultimate pinnules oblong, toothed; sori nearly mar- 



var. bipinnata Makino Fumoto-kaguma, Kujaku-fomoto- ginal, suborbicular; indusia hyaline, glabrous or with few se- 



shida. Closely allied to the preceding, evergreen, but tose hairs near base, undulate to entire. Shikoku, Kyushu; 



stouter; blades bipinnate, the rachis usually glabrous and rare. Formosa. 



smooth on upper side except near the top; pinnules sessile, 5. Microlepia yakusimensis Tagawa. M. obtusiloba 



oblique, ovate to oblong-ovate, 1-2 cm. long, 6-12 mm. wide, sensu auct. Japon., non Hayata; M. majuscula sensu auct. 



obtuse to subacute, toothed, often lobulate. ^Honshu (Awa Japon., non Moore Yakushima-kaguma. Rhizomes 



and Sagami Prov. and westw.), Shikoku, Kyushu; rare. creeping, about 5 mm. across, chestnut-brown, hairy; stipes 



3. Microlepia strigosa (Thunb.) Presl. Trichomanes 30-50 cm. long, scattered short-pubescent, glabrescent, fuscous- 

 strigosum Thunb.; Dic\sonia strigosa (Thunb.) Thunb.; Da- brown; blades thinly herbaceous, 30-50 cm. long, 25-30 cm. 

 vallia strigosa (Thunb.) Kunze; Dennstaedtia strigosa wide, subtripinnate, the rachis subglabrous, grooved above, 

 (Thunb.) J. Smith; Dickjonia japonica Sw.; M. japonica (Sw.) fuscous-brown, densely short-pubescent beneath; pinnae ovate- 



Presl Ishi-kaguma. Resembling the preceding in habit; deltoid, acuminate, petiolulate, the lower pinnae 15-20 cm. 



rhizomes short-creeping, firm, 3-5 mm. across, while young long, 5-10 cm. wide, pinnules pinnatiparted, long-acuminate 



with brownish multicellular jointed hairs; fronds evergreen, in the lower, acute to obtuse in the upper ones, hirsute be- 



50-150 cm. long, rather coarse, strigose; stipes 30-60 cm. long, neath on costas and veins, short-pubescent above on the costas; 



dull purple on the lower side, stramineous on the upper side, ultimate pinnules oblong, obtuse, oblique, crenately toothed, 



densely brownish hairy at base, sparsely so above, the callose sometimes lobulate; sori small; indusia pale, with a few hispid 



base of the hairs persistent; blades chartaceous-herbaceous, hairs on back. Kyushu; rare. 



dark green, paler beneath, broadly lanceolate, 40-100 cm. long. 



3. MONACHOSORUM Kunze Fuji-shida Zoku 



Terrestrial, with ascending short rhizomes covered at the tip by a mucous excretion; fronds thinly brown-scurfy while young; 

 stipes tufted; blades simply or 2- to 4-pinnate, often with bulbils in axils of pinnae or proliferous at the tip; ultimate pinnules 

 toothed or lobed, membranous, blackish when dried, glabrous except for sparse glandular short hairs on the axis, the lobes with 

 a single veinlet ending short of the tip; sori terminal or nearly so on the veinlets, intramarginal, rounded; indusia absent, the 

 receptacle scarcely raised; sporangia mixed with glandular hairs. About 5 species, Malaysia to Japan. 



