42 



Pteridaceae 



IB. Ultimate divisions of blades pectinately pinnatiparted, at least the terminal. 



6A. Costas and costules without spines at base of segments on upper side; veinlets reaching the awn-toothed sterile margin of the blade. 

 7A. Blades 40-60 cm. long, 20-25 cm. wide; stipes 4-angled, 3-4 mm. across near base; rachis rounded on back; stipes and rachis 



dark purple-brown, nearly concolorous; ultimate segments 5-10 mm. wide 6. P. semipinnata 



7B. Blades 25-40 cm. long, 8-15 cm. wide; stipes subtrapezoid or 3-angled, 2-2.5 mm. across near base; rachis 3-angled; stipes and 



rachis yellow-brown, with paler angles; ultimate segments 4-5 mm. wide 7. P. dispar 



6B. Ck)stas and costules with a spine at base of segments on upper side. 



8A. Veinlets reaching the margin of sterile part of blade; sterile segments entire 8. P. quadriaiirita 



8B. Veinlets not reaching the margin; sterile segments toothed. 



9A. Blades 20-30 cm. long, bipinnatiparted, the lowest pinnae sometimes with 1 or 2 sessile branches on posterior side near base. 



lOA. Pinnae gray- or brownish green; veinlets free 9. P. inaequalis 



lOB. Pinnae vivid green; lowest veinlets of the segments anastomosing, forming elongate areoles along the costas of the pinnules. 



10. P. nakflsimae 

 9B. Blades often more than 1 m. long, with 5 bipinnatiparted, short-stalked, large pinnae; lowest veinlets of the segments anastomos- 

 ing and forming areoles along the costas of the pinnules 11. P. wallichiana 



1. Pteris vittata L. P. longifolia sensu auct. Asiat., non 4. Pteris ryukyuensis Tagawa. Ryukyu-i-no-moto-s6. 



L. ^MoEjiMA-sHiDA. Rhizomcs short, rather stout, with Fronds dimorphic, glabrous, evergreen; rhizomes small, short, 



dense, pale brown, membranous, broadly linear scales about 3 with dark purple-brown, lustrous, broadly linear scales, about 



mm. long; stipes 3-15 cm. long, pale brown or stramineous, 1 mm. long; stipes firm, slender, obtusely angled, lustrous, 



sulcate on upper side, prominendy scaly, the callose base of brown, paler on the angles, 0.7-1 mm. across, 3-12 cm. long 



scales persistent; blades obovate-oblong to broadly oblanceolate, in the sterile fronds, 5-25 cm. long in the fertile; blades ternate 



20-50 cm. long inclusive of the elongate terminal pinna, nar- or sometimes with a pair of remote pinnae at base, the rachis 



rowed at base, simply pinnate, the rachis stramineous, sulcate not winged; sterile pinnae chartaceous to thinly coriaceous, 



on upper side, rounded and more or less scaly or scabrous on lanceolate to oblong-ovate, 2-7 (-10) cm. long, 7-12 mm. wide, 



underside; pinnae 12-26 pairs, spreading, simple, the lower obtuse, the lower ones sometimes 2-parted, the fertile pinnae 



ones 1-2 cm. long, the median 4-8 cm. long, truncate and linear, 4-15 cm. long, 3^ mm. wide. Stone walls; Kyushu 



sometimes slighdy auriculate at base, sessile, glabrous and (s. distr.) ; rare. Ryukyus. 



gray-green above, the costa thinly pubescent beneath, the 5. Pteris yamatensis (Tagawa) Tagawa. P. angusti- 



sterile pinnae linear-lanceolate to oblong-ovate, 7-8 mm. wide, pinna var. yamatensis Tagawa Hime-i-no-moto-s6. Some- 



usually obtuse, minutely toothed, the fertile pinnae elongate, what resembling the preceding in habit; rhizomes short, with 



4-7 mm. wide. Honshu (Kii Prov.), Shikoku, Kyushu (s. linear-lanceolate black-brown pale margined scales 2-2.5 mm. 



distr.); rare. Ryukyus, Formosa, s. China, and generally in long; fronds tufted, the sterile shorter than the fertile; stipes 



the Tropics and subtropics of the Old World. slender, trigonous, 0.5-1 mm. across, chesmut-brown, 2-8 cm. 



2. Pteris multifida Poir. P. serrulata L. f., non Forsk. long in the sterile fronds, about 12 cm. long in fertile; blades 



I-NO-MOTo-s6. Fronds slender, glabrous; rhizomes ternately or pedately 3- to 5-foliolate; sterile pinnae thinly 



short, with black-brown lustrous broadly linear entire scales chartaceous, linear, 3-13 cm. long, 3-6 mm. wide, the terminal 



about 2.5 mm. long; fronds tufted, the sterile ones shorter; pinna 10-20 cm. long, acutely toothed, gradually narrowed at 



stipes slender, pale green, 3-angled, glabrous in upper part, both ends, subsessile, glabrous but with scattered hairlike scales 



thinly scaly at base; blades broadly ovate in outline, the rachis along costas on both sides while young; veins simple or once 



winged; sterile pinnae thinly chartaceous, 2 or 3 on each side, forked, not reaching die margin; fertile pinnae 2-3 mm. wide, 



3- to 5-parted, the sterile divisions linear to lanceolate, 5-15 ejitire. Calcareous rocks in mountains; Honshu (Mount 



cm. long, 6-12 mm. wide, long-acuminate, sessile or decurrent Omine in Yamato Prov.) ; rare. 



on die rachis, irregularly aristulate-toothed; fertile pinnae lin- 6. Pteris semipinnata L. 0-Amakusa-shida. Stipes 



ear, entire, 10-20 cm. long, 3-7 mm. wide. Lowlands; Hon- 30-60 cm. long, 3-4 mm. across near base, lustrous, 4-angled, 



shu (Kanto Distr. and westw.), Shikoku, Kyushu; common. dark purple-brown or deep brown, smooth on upper part, scaly 

 China and Indochina. at base; blades 40-60 cm. long, 20-25 cm. wide, bipinnati- 



3. Pteris cretica L. P. nervosa Thunb. 5ba-no-i-no- parted, with a large terminal pinna, the rachis deep brown, 



MOTO-so. Fronds evergreen, glabrous, nearly all alike; rhi- lustrous, rounded on back, smoodi; lateral pinnae 5-8 pairs, 

 zomes short, widi dark purple-brown lustrous broadly linear 10-17 cm. long, subsessile, the long-acuminate slightly falcate 

 scales about 3 mm. long; stipes 15-60 cm. long, pale strami- tip 5-10 cm. long, unlobed and entire on anterior margin or 

 neous, sometimes somewhat brownish toward the base, ob- sometimes witii a small auricle at base, the terminal pinna reg- 

 tusely 4-angled, sulcate on the upper side, glabrous, 2-2.5 mm. ularly pinnatiparted, the costas with a sparsely scabrous line 

 across; blades thinly coriaceous, 20-40 cm. long, simply pin- above on each side; pinnules linear-lanceolate, about 8 cm long, 

 nate, with 1-5 pairs of pinnae, the rachis not winged; lower 5-10 mm. wide, acuminate, incurved-toothed on sterile mar- 

 pinnae usually 2- or 3-parted to -divided; fertile lobes broadly gins; costules sparsely scabrous on the upper side. Kyushu 



linear, 5-12 mm. wide, gradually long-acuminate, sterile at the (Yakushima) ; rare. Ryukyus, Formosa, China, and tropical 



tip, the sterile lobes linear-lanceolate, 10-25 cm. long, 1-2.5 cm. Asia. 



wide, acuminate, awn-toothed, green; veinlets reaching the 7. Pteris dispar Kunze. P. semipinnata var. dispar 



cartilaginous margin of the sterile pinnae. Honshu (Kanto (Kunze) Bak. Amakusa-shida. Fronds evergreen; rhi- 



Distr. and Echigo Prov. westw.), Shikoku, Kyushu; common. zomes short-creeping, with dark brown linear scales 3-3.5 mm. 



Widely distributed in the tropics and warm-temperate regions long; stipes tufted, 20-40 cm. long, yellow-brown, subtrapezoid 



of the N. Hemisphere. Cv. Albolineata. P. cretica var. or 3-angled, lustrous, 2-2.5 mm. across near base, smooth, ob- 



albolineata Hook. Matsuzaka-shida. Sterile lobes white- tuse on the dorsal angle; blades chartaceous, lanceolate to nar- 



variegated along the costas. Cultivated. rowly oblong-ovate, 25-40 cm. long, 8-15 cm. wide, bipinnati- 



