Pteridaceae 



45 



t^tthnii var. brandtii (Fr. & Sav.) Tagawa Miyama-urajiro. 



Rhizomes short, covered with stubs of old stipes and dark 

 brown lanceolate scales; fronds fragile, tufted; stipes 3-15 cm. 

 long, terete, purple- to red-brown, lustrous, the scales at base 

 membranous, lanceolate, 4-6 mm. long, finely striate; blades 

 usually longer than the stipes, ovate to oblong-ovate, 8-30 cm. 

 long, 4-12 cm. wide, glabrous on upper side, more or less 

 white-farinose beneath, 2- or 3-pinnatiparted, acuminate, acute 

 at the tip, the rachis purplish, sulcate on upper side; pinnae 



sessile; pinnules oblong-ovate to oblong, obtuse to subacute, 

 lobed to parted, the lower ones usually sessile, the upper ones 

 and sometimes the lower pinnules decurrent on the rachis, 

 crenate; veins very slender, usually invisible on underside; 



sori marginal; false indusia lobulate, scarious. Mountains; 



Honshu (centr. and Kanto Distr.); rather rare. Forma 



efarinosa (Makino) Tagawa. Cheilanthes brandtii var. ejar- 



inosa Makino. Blades green beneath. The typical phase 



occurs in Korea, China, Manchuria, and Amur. 



13. CRYPTOGRAMMA R. BR. 



RiSHIRI-SHINOBU ZoKU 



Terrestrial with solenostelic or dictyostelic rhizomes; fronds tufted or remote, small, herbaceous, glabrous, pinnately dis- 

 sected, often dimorphic, the fertile fronds with narrower and longer pinnules than the sterile; veins free; sori submarginal, on 



the branches of the forked veins, covered by a mosdy continuous reflexed margin (false indusium) ; paraphyses absent. ^Four 



species, widely distributed in cold regions of the N. Hemisphere. 



I A. Rhizomes rather short, densely leafy and thickly clothed with stubs of old stipes; fronds tufted; blades 2- to 4-pinnate, herbaceous. 



1. C. crispa 

 IB. Rhizomes very slender, sparsely leafy, without old stipes; fronds not tufted; blades 1- or 2-pinnate, membranous 2. C. stelleri 



1. Crj'ptogramma crispa (L.) R. Br. Osmunda crispa 



L.; Pteris crispa (L.) AH.; Allosorus aisptis (L.) Bernh. 



RisHiRi-SHiNOBU. Rhizomes rather short, ascending, densely 

 clothed with stubs of old stipes; fronds glabrous, tufted; stipes 

 4-20 cm. long, light brown, 1.5-2 mm. across at base, smooth, 

 sulcate on upper side, sparsely scaly at base; scales on stipes and 

 rhizomes membranous, broadly lanceolate to narrowly deltoid- 

 ovate, 4-6 mm. long, pale brown, deciduous; blades herba- 

 ceous, ovate to narrowly so, 5-15 cm. long, 2.5-6 cm. wide, 

 acute, yellowish green, 2- to 4-pinnate; pinnae 4-7 on each side, 

 oblong-ovate, short-petiolate; ultimate sterile pinnules ovate to 

 oblong, 5-8 mm. long, 2-6 mm. wide, few-toothed or lobed to 

 parted; veinlets not reaching the margin of the tooth; ultimate 

 fertile pinnules lanceolate, 4-10 mm. long, 2-4 mm. wide, sub- 

 obtuse, crenate; sori terminal on the veinlets; false indusium 

 hyaline, undulate. July-Oct. Rocky alpine slopes; Hok- 

 kaido, Honshu (n. distr.); rare. s. Kuriles, s. Sakhalin, 



Siberia to Europe. 



2. Cryptogramma stelleri (Gmel.) Prand. Pteris stel- 

 leri Gmel.; Allosorus stelleri (Gmel.) R. Br.; Pellaea stelleri 



(Gmel.) Bak.; Pteris gracilis Michx. Yatsuga-take-shinobu. 



Rhizomes creeping, very slender, about 1.5 mm. across, sparsely 

 leafy and scaly; scales broadly lanceolate, 1-2 mm. long, fuga- 

 cious, pale brown; fronds delicate, glabrous; stipes 3-10 cm. 

 long, 0.5-0.7 mm. across, light brown, slighdy lustrous, 

 smooth, sparsely scaly at base; blades membranous, oblong- 

 ovate or ovate, 3-7 cm. long, 2-3 cm. wide, 1- or 2-pinnate; 

 pinnae simple or ternate, very short-petiolulate to subsessUe; 

 sterile pinnules ovate to obovate or flabellate, 4-7 mm. long, 

 rounded to crenate-toothed on upper half, broadly cuneate and 

 entire on lower portion; fertile lobes broadly lanceolate, 5-12 



mm. long, 3-4 mm. wide, crenate to subentire. July-Sept. 



Wet shaded rocks, alpine regions; Honshu (centr. distr.); very 

 rare. Siberia, the Himalayas, and N. America. 



14. ONYCfflUM Kaulf. Tachi-shinobu Zoku 



Terrestrial; rhizomes creeping, or short and compact, solenostelic, scaly; fronds tripinnate to decompound; pinnules small, 

 herbaceous to subcoriaceous, narrow, glabrous; veins free except the fertile commissure connecting the tips; sori continuous 



along both margins, covered by the scarious reflexed marginal or submarginal false indusium; paraphyses absent. Few species, 



in die Old World. 



1. Onychium japonicum (Thunb.) Kunze. Tricho- 

 nianes japonicum Thunb.; Caenopteris japonica (Thunb.) 

 Thunb.; Cryptogramma japonica (Thunb.) Prand Tachi- 

 shinobu, Kan-shinobu. Rhizomes creeping, sparsely leafy, 

 2.5-4 mm. across, with brown membranous broadly lanceolate 

 scales 2.5-3 mm. long; fronds evergreen, glabrous; stipes slen- 

 der, 10-50 cm. long, 1.5-3 mm. across near base, pale green to 

 straw-colored, smooth, striate on the upper side, purple-brown 

 toward the base; blades triangular-ovate, 15-40 cm. long, 8-20 

 cm. wide, acuminate, deep green on upper side, paler beneath, 

 3- or 4-plnnate; pinnae obliquely ascending, gradually smaller 



in the upper ones, alternate, the lowest ones 1/2-2/3 as long 

 as the blade, triangular-ovate, long-acuminate, prominendy 

 petiolulate; pinnules narrowly lanceolate, 5-8 mm. long, 1-1.5 

 mm. wide, acute, entire, the sterile ones often few-toothed; 

 veins unbranched in the sterile pinnules, pinnately branched 

 in fertile ones; sori marginal; false indusia hyaline, paired, 



often overlapping while young, entire, 3-8 mm. long. Low 



mountains; Honshu (Kanto Distr. and westw.), Shikoku, 

 Kyushu; rather common. s. Korea, China, Formosa, Malay- 

 sia to India. 



15. PLEUROSORIOPSIS Fomin Karakusa shida Zoku 



Terrestrial rhizomatous fern; fronds remote, small; blades ovate to ovate-oblong, herbaceous, l(-2)-pinnate, with numerous 

 spreading brown multicellular hairs; veins free, not prominent on both sides, not reaching the margin of the pinnules; sporangia 

 along the lower and middle portions of the veins, naked. A single species. 



