64 ASPIDIACEAE 



HA. Pinnae more than 10 pairs. 



15A. Pinnae slightly dilated at base; fronds gradually narrowed at base; sori along the costa, usually in 2 series; indusium 



1.2-1.5 mm. across 12. D. tokyoensis 



15B. Pinnae not dilated at base; fronds slightly or not narrowed at base; sori between the costa and margin, in irregular 

 series; indusium about 0.5 mm. across. 



16A. Sori nearer the margin than the costa; scales on stipes and rachis brown 13. D. dicl^insii 



16B. Sori nearer the costa than the margin; scales on stipes and rachis black-brown to purplish brown 14. Z). atrata 



14B. Pinnae 2-7 pairs. 



17A. Pinnae 2-5 pairs, unlobed 15. D. sieboldii 



17B. Pinnae 5-7 pairs, pinnately parted in the lower portion, lobed in the upper portion; terminal pinnae pinnatiparted. 



16. D. loyamae 

 4B, Scales on blades nearly absent. 



18A. Stipes densely scaly on lower half, the basal scales ovate, abruptly acuminate; axis of pinnae winged in upper portion. 



17. D. sabae 

 18B. Stipes sparsely scaly throughout, the basal scales lanceolate, gradually acuminate; axis of pinnae more or less winged through- 

 out. 

 19A. Basal scales chartaceous, 6-10 mm. long; indusium strongly incurved on margin, at maturity more or less 2- or 3-lobed. 



18. D. hayatae 

 19B. Basal scales membranous, 10-15 mm. long; indusium slightly incurved on margin, not breaking apart at maturity. 



19. D. sparsa 

 3B. Scales on blades beneath and on upper portion of stipes prominently dilated or saccate, nearly absent in Nos. 30 and 31. 



20A. Blades simply pinnate 20. D. decipiens 



20B. Blades bi- to tri (-4) -pinnate. 



21A. Lowermost posterior pinnules of lowest pinnae not larger than the adjacent ones. 



22A. Scales of stipes and rachis linear- to lanceolate-deltoid, prominently toothed, brown 21. D. championii 



22B. Scales of stipes and rachis scattered, linear to linear-lanceolate, entire or rarely obsoletely toothed, brown to black-brown. 

 23A. Pinnules entire or crenately mucronate-toothed, broadly lanceolate, elliptic to oblong, acute to obtuse or truncate at apex. 



22. D. fuscipes 

 23B. Pinnules coarsely toothed to pinnatiparted, lanceolate, linear-lanceolate, or oblong. 



24A. Blades deltoid to deltoid-ovate; pinnae and pinnules horizontally spreading; pinnules deltoid-ovate to narrowly oblong, 



at least the lower ones pinnatiparted 23. D. indusiata 



24B. Blades oblong to ovate-oblong; pinnae and pinnules oblique to nearly horizontally spreading; pinnules linear-lanceolate, 

 oblong, or oblong-ovate. 



25A. Pinnae sessile or nearly so 24. D. erythrosora 



25B. Pinnae distinctly petiolulate 25. Z). hondoensis 



2 IB. Lowermost posterior pinnules of lowest pinnae larger than the adjacent ones. 

 26A. Indusium present. 



27A. Blades ovate to oblong-ovate, much longer than wide; lowest pinnae mostly less than half as long as the entire blade; 

 scales on stipes and rachis linear to linear-lanceolate. 

 2 8 A. Scales on rachis and stipes appressed to ascending. 



29A. Scales on stipes and rachis ascending, mostly black-brown at base, rarely brown and concolorous 26. D. varia 



29B. Scales on stipes and rachis appressed, nearly uniformly black, concolorous 27. D. sordidipes 



28B. Scales on rachis and stipes spreading 28. D. saxifraga 



27B. Blades 5-angled, or deltoid with an angle at base on each side, nearly as wide as long; lowest pinnae mostly more than 

 half as long as the entire blade, rarely less, if so, then the scales on stipes broadly lanceolate. 

 30A. Scales on rachis and on underside of secondary axis rather dense, subulate-linear, black-brown, saccate at base; lowest 



pinnae subsessile to very short-petiolate; pinnules with incurved short awn-tipped teeth 29. D. formosana 



30B. Scales on rachis and secondary axis very scanty or nearly absent; lowest pinnae distinctly petiolulate; pinnules with 

 ascending acute teeth or mucronate-toothed. 

 31 A. Scales thinly scattered beneath on costules of pinnules, stipes, and rachis; pinnae thinly herbaceous, the lowest with 



petiolules 1—2.5 cm. long 30. D. chinensis 



31B. Plant glabrate; pinnae thickly herbaceous, the lowest with petiolules 2-5 cm. long 31. D. gymnophylla 



26B. Indusium absent 32. D. gymnosora 



1. Dryopteris sikokiana (Makino) C. Chr. Ctenitis nearly glabrous on both sides, sometimes with a fevs' scattered 



si\o\iana (Makino) H. Ito; Nephrodium sik_okianum Ma- fleshy hairs on upper side, the costules above with an elevated 



kino HoNOKAwA-sHiDA. Rhizomes short-creeping; stipes ridge on each side, the lowest pinnule largest, sometimes bipin- 



20-50 cm. long, brownish, densely scaly; scales on lower half natiparted; ultimate pinnules oblong to oblong-lanceolate, 5-20 



of the stipes broadly lanceolate to narrowly ovate, 7-10 mm. mm. long, 2-7 mm. wide, obtuse, crenate; sori nearer the costa 



long, entire, dark chestnut-brown, lustrous, spreading to (or to the veins) of the pinnules; indusia obsolete or absent. 



slightly deflexed, those on upper half of stipe and those of the Honshu (Izu, Echizen, and Kii Prov.), Shikoku, Kyushu. 



rachis linear to linear-lanceolate, 3-8 mm. long; blades ovate, 2. Dryopteris hendersonii (Bedd.) C. Chr. Ctenitis 



40-75 cm. long, acuminate, 3-pinnatiparted; pinnae 3-10 pairs, hendersonii (Bedd.) H. Ito; Lastrea hendersonii Bedd.; Neph- 



narrowly oblong-ovate to broadly lanceolate, equilateral, ob- rodium spectabile C. B. Clarke; L. spectabilis (C. B. Clarke) 



liquely spreading, caudate-acuminate, short-petiolulate; pin- Bedd.; D. leptorhachia Hayata Horai-hime-warabi. Rhi- 



nules narrowly oblong, obtuse in the smaller ones, narrowly zomes short-creeping; stipes 40-60(-100) cm. long, deep 



ovate-oblong, 2-6 cm. long, 1-2.5 cm. wide, short-acuminate in brown, 3-6 mm. across, scaly; scales spreading to reflexcd, 



larger ones, pinnatilobed, sometimes slightly whitish beneath, brown, lanceolate to linear-lanceolate, 5-15 mm. long, entire; 



