76 



ASPIDIACEAE 



8-12 cm. wide, acute, cordate, jointed with the stipe, pinnati- 

 parted to -cleft, smooth; pinnules 7-12 pairs, oblong-lanceo- 

 late, 1-1.5 cm. wide, horizontally spreading, obtuse to sub- 

 acute, pinnately lobed, crenate-toothed, the lowest pinnules 

 broadly lanceolate, to 2 cm. wide, more or less deflexed, 

 slightly broadened on the posterior side; veins pinnately 



branched, the veinlets simple or once forked; sori elliptic, 1-2 

 mm. long, sometimes oblong or rounded, in 2 series on the 

 nearer pinnules, the costa in several somewhat irregular series. 



Honshu (Kanto to s. Kinki distr.). Shikoku. China 



(Philippines to New Guinea?). 



15. CYCLOSORUS Link Ho-shida Zoku 



Rhizomes erect to creeping; blades bipinnatiparted; veins pinnately branched, the single or sometimes few lower veinlets 

 united with those of the adjacent veins at or below the sinus; hairs short, acicular and glandless or glandular, often mixed to- 

 gether; sori on the veinlets rounded; indusia orbicular-reniform, usually large, the sporangia often with few, simple or 

 glandular hairs. About 300 species, in the Tropics and subtropics. 



lA. Lowest veinlet only fused to the adjacent vein; blades regularly bipinnatiparted. 



2A. Pinnae coriaceous to firmly chartaceous, with small ovate scales on costas beneath; pinnules broadly deltoid-ovate, mucronate; vein- 

 lets very close; sori marginal; seaside plant 1. C. goggilodus 



2B. Pinnae chartaceous, not scaly; pinnules ovate to oblong, acute to very obtuse, sometimes mucronate; veinlets not very close; sori 

 median, or nearer the margin than the costa. 

 3A. Stipes tufted on short ascending rhizomes; blades narrowed at base; lowest anterior segment not obviously larger than the others; 



scales puberulous 2. C. dentatus 



3B. Stipes distant on long-creeping rhizomes; blades not at all narrowed at base; lowest anterior segment larger than the others; scales 

 glabrous. 



4 A. Pinnae sparsely hairy, often glabrescent above except for the costas 3. C. acuminatus 



4B. Pinnae prominently hairy on both sides 4. C. parasiticus 



IB. Nearly all of the veinlets fused to those of the adjacent veins; blades simple or with a few unlobed pinnae 5. C. triphyllus 



1. Cyclosorus goggilodus (Schk.) Link. Aspidium 

 goggilodus Schk.; Dryopteris goggiloda (Schk.) O. Kuntze; 



Dryopteris unita sensu auct. Japon., non O. Kuntze Tetsu- 



Ho-SHIDA. Plant nearly glabrous; rhizomes long-creeping, 

 with scattered brown ovate scales 2-3 mm. long; stipes 30-90 

 cm. long, 3-4 mm. across, lustrous, brownish, nearly naked; 

 blades lanceolate to narrowly oblong, 30-70 cm. long, 10-25 

 cm. wide, abruptly contracted at tip; pinnae 20-28 pairs, 

 coriaceous to firmly chartaceous, broadly linear, 8-15 mm. 

 wide, acuminate, cuneate at base, subsessile, subglabrous to 

 short-pilose beneath, nearly glabrous above, with small ovate 

 scales 0.5-1 mm. long on costas beneath, the terminal pinna 

 similar to the lateral ones; segments broadly deltoid-ovate, 

 mucronate; veins 6-8 pairs per segment, very close; sori sub- 

 marginal; indusia orbicular-reniform, 0.2-0.5 mm. across, 



nearly glabrous to short-pilose. Wet places near the sea; 



Honshu (Izu Prov. and westw. in warmer parts), Shikoku, 



Kyushu. ^Ryukyus, Formosa, and China; widely distributed 



in the tropics and subtropics. 



2. Cyclosorus dentatus (Forsk.) Ching. Polypodium 

 dentatum Forsk.; C. oblancifolius (Tagawa) Tagawa; Dryo- 

 pteris oblancijolia Tagawa Inu-keho-shida. Rhizomes 



short, ascending; stipes tufted, short-pilose, sparsely scaly; 

 scales dark brown, linear, about 10 mm. long, puberulous; 

 blades herbaceous, oblong-lanceolate to oblanceolate-oblong, 

 30-60 cm. long, 10-16 cm. wide, gradually narrowed at base, 

 rather abruptly narrowed toward tip, the terminal pinna 

 deltoid-lanceolate, 10-12 cm. long, 2-3 cm. wide; rachis densely 

 pubescent on upper side, less so beneath; pinnae 13-15 pairs, 

 spreading, broadly linear-lanceolate, 6-8 cm. long, 10-17 mm. 

 wide, abruptly acuminate, truncate at base, sessile, pinnately 

 cleft, short-pilose on both sides, densely long-pilose on the 

 costas, sparsely so on the veinlets, the lower pinnae reduced, 

 ovate, 1,5-2 cm. long, about 1 cm. wide, obtuse; pinnules 

 oblong, about 3 mm. wide, obtuse or truncate-rounded at 

 apex, entire; veinlets simple, not reaching the margin; sori 

 median; indusia orbicular-reniform, densely short-pilose. 



Honshu (Kii Prov.), Shikoku, Kyushu (s. distr.) ; rare. 



Formosa and widespread in the Tropics. 



3. Cyclosorus acuminatus (Houtt.) Nakai. Polypod- 

 ium acuminatum Houtt.; Dryopteris acuminata (Houtt.) 

 Nakai; Polypodium sophoroides Thunb.; Aspidium sopho- 

 roides (Thunb.) Sw.; Nephrodium sophoroides (Thunb.) 

 Desv.; D. sophoroides (Thunb.) O. Kuntze; C. sophoroides 

 (Thunb.) Tard.-Bl.; A. oshimense Christ; D. oshimensis 

 (Christ) C. Chr.; D. ogatana Koidz.; D. ensipinna Tagawa 



Ho-SHiDA. Evergreen; rhizomes long-creeping, 3-4 



mm. across, sparsely pilose and thinly scaly; stipes slender, 

 25-50 cm. long, straw-colored, sometimes dull purplish, 

 sparsely short-pilose, sparsely scaly at base; scales lanceolate, 

 4-6 mm. long, brown; blades chartaceous, lanceolate to nar- 

 rowly oblong-ovate, 30-60 cm. long, pinnate, deep green above, 

 slightly paler beneath, abruptly and caudately long-acuminate, 

 the terminal pinna linear- to deltoid-lanceolate, the rachis 

 prominently short-pilose, somewhat hispid on underside; pin- 

 nae linear- to broad-lanceolate, 7-20 mm. wide, gradually 

 acuminate, spreading, truncate at base, sessile, pinnately cleft, 

 costas and veins strigose on upper side, spreadlng-pilose be- 

 neath; pinnules ovate to oblong-ovate, 3-4 mm. wide, obtuse 

 to subacute with a short mucro at tip, the lowest anterior 

 pinnule longer than the others; sori in 2 rows, nearer the 

 margin than the veins; indusia small, orbicular-reniform, 

 short-pilose on back, about 0.5 mm. across, entire, brownish. 



Thickets and sunny slopes; Honshu (Kanto Distr. 



through Tokaido to Kinki and westw.), Shikoku, Kyushu. 

 Ryukyus, Formosa, s. Korea, and China. 



4. Cyclosorus parasiticus (L.) Farw. Polypodium 

 parasiticum L.; Aspidium parasiticum L.; Nephrodium para- 



siticum (L.) Desv.; Dryopteris parasitica (L.) O. Kuntze 



Ke-ho-shida. Rhizomes long-creeping; stipes 20-80 cm. long, 

 pilose, scaly toward the base; scales dark brown, linear to 

 linear-lanceolate, 8-15 mm. long, entire; blades chartaceous, 

 oblong-lanceolate, 40-70 cm. long, 15-30 cm. wide, abrupdy 

 contracted and acuminate at the tip, prominently pubescent on 



