ASPIDIACEAE 



85 



gawa; A. yo\oscensc var. alpicola Hiyama Hebi-no-negoza. 



Rhizomes short, erect, covered with the sti.ibs of old stipes; 

 fronds green, glabrous, somewhat tufted; stipes 15-25 cm. long, 

 straw-colored, scaly toward base; scales membranous, linear to 

 linear-lanceolate, 8-15 mm. long, about 2 mm. wide, uniformly 

 brown or with a darker somewhat lustrous thicker band in 

 the center, entire; blades herbaceous, oblong-ovate, 25-40 cm. 

 long, 8-20 cm. wide, acuminate, scarcely narrowed at base, 

 sessile, 2-pinnate; pinnae 10-15 on each side, equilateral, 

 lanceolate, 1-2.5 cm. wide, long-acuminate, the rachis scarcely 

 winged; pinnules oblong-ovate, 5-12 mm. long, 3-5 mm. wide, 

 obtuse to acute, toothed to lobulate, very slightly adnate to 

 tlie rachis, scarcely decurrent; sori 4-10 to a pinnule, median; 

 indusia about 1 mm. long, oblong, straight, the lower ones 



often hamate (athyrioid), thin, entire. Woods in lowlands 



and mountains; Hokkaido, Honshu, Shikoku, Kyushu; com- 

 mon and variable. Sakhalin, Korea, Manchuria, and e. 



Siberia. 



Var. fauriei (Christ) Tagawa. Nep/irodium jattrici Christ; 

 A. fauriei (Christ) Makino; A. niWoense Makino; Aspidium 

 fauriei (Christ) Christ; Athyrium demissum Christ, pro parte 



IwA-iNU-w.-\RABi. Depauperate, with smaller, narrower 



bipinnatifid blades narrowed at base, the pinnae acute to 



obtuse. Rocky places in mountains; occurs with the typical 



phase. s. Kuriles and Korea. 



Poorly known are: Athyrium kirishimense Tagawa. 

 KiRiSHiM.\-HEBi-No-NEGoZA. Reported to have larger scales, 

 to 17 mm. long at base of stipes, and chartaceous-herbaceous 



blades. Kyushu (centr. distr.). Athyrium satowii H. 



Ito ToRA-No-o-iNU-WARABi. An aberrant phase of A. 



yo\osccnse with the lower pinnae broadly Unear, rounded, 

 pinnatiparted, the upper pinnae short, simple, flabellate, the 

 pinnules rounded-obovate, the sori few near die margin. 



29. Athyrium tashiroi Tagawa. Usuea-hebi-no-negoza. 

 Rhizomes erect or ascending, covered with the basal stubs of 

 old fronds; stipes tufted, 15-25 cm. long, slender, straw- 

 colored, dark brown and densely scaly toward base; scales 

 membranous, narrowly lanceolate to linear, 1-1.5 cm. long, 

 to 2 mm. wide, entire, uniformly brown or sometimes with 

 a chestnut-brown longitudinal band in the center; blades 

 herbaceous, ovate, 15-30 cm. long, 10-18 cm. wide, acuminate, 

 bipinnate, glabrous; pinnae 10-15 pairs, spreading, broadly 

 ovate-lanceolate, 8-20 cm. long, 2-3.4 cm. wide, caudately 

 long-acuminate, sessile; pinnules several to rather many on a 

 pinna, rather remote, broadly lanceolate to narrowly ovate, 

 1.5-2 cm. long, 4-8 mm. wide, acute to short-acuminate, 

 oblique, cuneate at base, sessile, irregularly few-toothed, pin- 

 natcly cleft to parted, pinnules ascending, acuminate; sori 

 median; indusia membranous, athyrioid, sometimes straight, 

 entire, about 1 mm. long. Mountains; Kyushu; rare. Pos- 

 sibly only a variety of A. yo\oscense. 



30. Athyrium multifidum Rosenst. A. filix-fcmitia var. 

 deltoideuin Makino; A. sohitum Rosenst., non Christ; A. 

 multifidum var. latisectum Rosenst.; A. deltoidofrons Ma- 

 kino; A. multifidum var. deltoideum (Makino) Nakai; A. 

 deltoidofrons var. multifidum (Rosenst.) Koidz. and var. 



latisectum (Rosenst.) Koidz. Sato-me-shida, O-sato-me- 



SHiDA. Fronds large, glabrous, tufted; stipes rather stout, 40- 

 60 cm. long, straw-colored, densely scaly near base; scales mem- 

 branous, pale brown, linear-lanceolate to lanceolate, 10-15 

 mm. long, 1-2 mm. wide, long-acuminate, entire; blades thinly 

 herbaceous, deltoid-ovate to broadly ovate, 40-70 cm. long. 



30-50 cm. wide, acuminate, 3-pinnatiparted; pinnae 9-12 

 pairs, equilateral, spreading, 15-30 cm. long, 4-7 cm. wide, 

 gradually acuminate, short-petiolulate; primary pinnules nar- 

 rowly deltoid-ovate to deltoid-lanceolate, 2.5-4 cm. long, 8-12 

 mm. wide, acute to acuminate, very short-petiolulate to sub- 

 sessile, pinnatiparted nearly to the costas; ultimate pinnules ob- 

 long to ovate, 4-7 mm. long, obtuse to subacute, toothed; sori 

 5-10 to a pinnule, subcostal; indusia thin, oblong and straight, 

 0.7-1 mm. long, sometimes athyrioid, erosc-ciliate. Hok- 

 kaido, Honshu, Shikoku. 



Var. acutissimum (Kodama) Ohwi. A. acutissimum Ko- 

 dama; A. deltoidofrons var. acutissimum (Koidz.) Tagawa; 



A. solutum var. acutissimum (Kodama) Tagawa Togariba- 



me-shida. Plant smaller, with narrower blades, and the 

 pinnae often horizontally spreading, usually 10-20 cm. long, 

 and with less deeply parted pinnules. Honshu and Shikoku. 



31. Athyrium pinetorum Tagawa. Takane-sato-me- 

 SHiDA. Rhizomes erect or nearly so, covered with the stubs 

 of old stipes; stipes 15-30 cm. long, slender, straw-colored, 

 dark brown toward base, thinly scaly in upper part, more 

 densely so near base; scales near base of stipe narrowly lanceo- 

 late, 10-15 mm. long, 1.5-2 mm. wide, entire, slightly lustrous, 

 with a dark brown longitudinal band in the center, the scales 

 on upper part of the stipes shorter and thinner, lighter brown, 

 often curled, lanceolate, 7-10 mm. long, 2-3 mm. wide; 

 blades thinly herbaceous, ovate-deltoid to nearly deltoid, 17-30 

 cm. long, 15-25 cm. wide, acuminate, not narrowed at base, 

 bipinnate or tripinnatiparted, sparsely scaly while young, 

 glabrescent; pinnae 7-14 cm. long, 3-4.5 cm. wide, acuminate, 

 equilateral, slighdy narrowed at base, sessile; pinnules nar- 

 rowly oblong-ovate, 15-20 mm. long, 5-9 mm. wide, obtuse to 

 subacute, slighdy oblique, sessile, pinnately cleft to parted; 

 pinnules elliptic, 2-4 mm. long, few-toothed; sori 7-12 to a 

 pinnule, subcostal; indusia thin, oblong, straight, often athyri- 

 oid, about 1 mm. long, erose-ciliate. Mountains; Hok- 

 kaido, Honshu (centr. distr. to Kii Prov.), Shikoku; rather 

 rare. 



32. Athyrium otophorum (Miq.) Koidz. Asplenium 

 otophorum Miq.; Diplazium otophorurn (Miq.) C. Chr.; 



Athyrium rigescens Makino Tani-inu-warabi. Rhizomes 



erect, short, covered with the basal stubs of old fronds; fronds 

 tufted; stipes 20-30 cm. long, straw-colored to pale brown, 

 usually slighdy purplish, densely scaly at base; scales broadly 

 linear, 8-10 mm. long, about 1 mm. wide, nearly black, 

 slighdy lustrous, entire; blades firmly herbaceous, broadly 

 ovate to oblong-ovate, 25-45 cm. long, 15-30 cm. wide, 

 abrupdy caudate-acuminate, not narrowed at base, glabrous, 

 bipinnate; pinnae 8-10 pairs, lanceolate, 2-4 cm. wide, long- 

 acuminate, spreading, sessile or nearly so; pinnules rather 

 numerous, the basal one auriculate on anterior side, narrowly 

 ovate, 1.2-2.5 cm. long, 5-12 mm. wide, acute to obtuse, 

 broadly cuneate at base, sessile, toothed to lobulate; sori 8-14 

 to a pinnule, subcostal, parallel, ascending; indusia broadly 

 lanceolate, subcntire, often brownish, straight to gently curved, 



1.5-4 mm. long, the lowest anterior one often athyrioid. 



Honshu (Tokaido to Kinki Distr. and westw.), Shikoku, 

 Kyushu. China. 



Athyrium yakusimense Tagawa, of Kyushu (Yakushima), 

 with paler scales, broader blades, and scarcely developed fleshy 

 bristles on the rachis of pinnae above at the insertion of pin- 

 nules, may be a form of A. otophorum. 



