ASPIDIACEAE 



33. Athyrium subrigescens (Hayata) Hayata ex H. Ito. 

 Diplazium subrigescens Hayata; A. elegans Tagawa; A. ele- 



gans var. purpurascens Tagawa Otome-inu-warabi, Horai- 



iNu-WARABi. Rhizomes short, erect; fronds deep green, gla- 

 brous, tufted; stipes straw-colored, 15-30 cm. long, densely 

 scaly toward the base; scales at base of stipes broadly linear, 

 8-12 mm. long, 1-2 mm. wide, black- to chestnut-brown, 

 usually with a paler margin, entire, slightly lustrous; scales 

 on upper part of stipe very sparse, membranous, early decidu- 

 ous, linear, 3-5 mm. long, pale brown; blades oblong and 

 narrowly long-acuminate, or ovate-oblong, 20-45 cm. long, 

 12-20 cm. wide, not or scarcely narrowed at base, bipinnate; 

 pinnae 10-15 pairs, equilateral, broadly linear to linear-lanceo- 

 late, 7-15 cm. long, 15-25 mm. wide, gradually acuminate, 

 falcate, sessile or nearly so; pinnules ovate, 8-12 mm. long, 

 rounded to obtuse, obliquely cuneate at base, subsessile, toothed 

 to lobulate; sori 3-10 to a pinnule, costal, ascending; indusia 

 broadly lanceolate, 1-2 mm. long, straight, rarely diplazioid. 

 ^Mountains; Kyushu (Yakushima); rare. ^Formosa. 



34. Athyrium niponicum (Mett.) Hance. Asplenium 

 niponicum Mett.; Asplenium uropteron Miq.; Athyrium uro- 



pteron (Miq.) C. Chr.; A. matsumurae Christ Inu-warabi. 



Plant glabrous; rhizomes slender, creeping, rather firm, 

 about 3 mm. across, scaly; stipes 20-40 cm. long, straw- 

 colored, very sparingly scaly; scales pale brown, membranous, 

 broadly linear, 4-6 mm. long, about 1 mm. wide, deciduous; 

 blades herbaceous, ovate to narrowly so, 25-35 cm. long, 12-25 

 cm. wide, acuminate-caudate, usually bipinnate; pinnae 6-10 

 pairs, alternate except the lowest 2, broadly lanceolate, 3-6 cm. 

 wide, long-acuminate, somewhat oblique, petiolulate; pinnules 

 broadly lanceolate to ovate, 1-3 cm. long, 3-12 mm. wide, 

 obtuse to acuminate, pinnately parted to lobulate, acutely 

 toothed, the lower ones often very short-petiolulate, the upper 

 pinnules decurrent on the rachis; sori on the ultimate pinnules, 

 linear, closely parallel; indusia lunate-lanceolate, 1-1.5 mm. 



long, frequently hamate (athyrioid), irregularly dentate. 



Shaded places in lowlands; Hokkaido (sw. distr.), Honshu, 



Shikoku, Kyushu; common. Korea, Manchuria, China, and 



Formosa. 



35. Athyrium vidalii (Fr. & Sav.) Nakai. Asplenium 

 vidalii Fr. & Sav.; Athyrium vidalii var. confusum Miyabe & 

 Kudo; A. commixtum Koidz. ^Yama-inu-warabi. Rhi- 

 zomes short, erect or ascending, densely covered with the basal 

 stubs of old stipes; stipes 20-50 cm. long, straw-colored, densely 

 scaly near base; basal scales broadly linear, 8-12 mm. long, 

 about 1 mm. wide, entire, brown; blades ovate to deltoid-ovate, 

 25-50 cm. long, 20-40 cm. wide, acuminate, not or scarcely 

 narrowed at base, bipinnate; pinnae broadly lanceolate, 3-5 

 cm. wide, long-acuminate, equilateral, short-petiolulate or 

 sometimes subsessile; pinnules sessile or very short-petiolulate, 

 oblique, deltoid-ovate to -lanceolate, 15-30 mm. long, 6-13 

 mm. wide, acute to acuminate, toothed, cuneate to subtrun- 

 cate at base, pinnately lobed to parted, the ultimate pinnules 

 elliptic to oblong, rounded; sori subcostal, 8-15 to a pinnule, 

 lanceolate to linear; indusia lanceolate, 1-2 mm. long, ascend- 

 ing, subentire, straight or hamate. ^Woods in mountains; 



Hokkaido, Honshu, Shikoku, Kyushu; rather conunon and 

 variable. Korea and Formosa. 



Var. yamadae (Miyabe & Kudo) Miyabe & Tatew. Ezo- 

 iNU-WARABi. Pinnules pinnately parted; indusia strongly 

 erose-toothed. Hokkaido. 



Athyrium arisanense (Hayata) Tagawa. Diplazium arisa- 



nense Hayata. A Formosan species with smaller fronds, obtuse 

 pinnules and median sori. Reported from Kyushu (Yaku- 

 shima). 



36. Athyrium wardii (Hook.) Makino. Asplenium 



wardii Hook.; Athyrium tsusimense Koidz. Hiroha-no- 



iNu-WARABi. Rhizomes short, erect, densely covered with 

 the basal stubs of old stipes; stipes 10-20 cm. long, straw- 

 colored, densely scaly near base; scales broadly linear, 6-13 

 mm. long, about 1 mm. wide, long-acuminate, with a dark 

 chestnut-brown longitudinal band down the center; blades 

 firmly herbaceous, bipinnate, narrowly to broadly ovate, 20-35 

 cm. long, 20-30 cm. wide, abruptly long-acuminate with the 

 tip more than one-third as long as the blade; pinnae usually 

 4-6 pairs, broadly lanceolate, 2.5-4 cm. wide, long-acuminate, 

 the lower ones short-petiolulate, not contracted at base except 

 the lowest one; pinnules elliptic to narrowly ovate, 1.5-3.5 cm. 

 long, 6-10 mm. wide, oblique, the lower ones broadly cuneate 

 at base, sessile, the upper ones more or less adnate and slightly 

 decurrent on the rachis, toothed to pinnately lobulate; sori 

 6-14 on each pinnule, nearer the costules than the margin; in- 

 dusia linear to lanceolate, 2-3.5 mm. long, straight, subentire, 

 the lowest anterior one rarely diplazioid. Woods in moun- 

 tains; Hokkaido, Honshu, Shikoku, Kyushu. Korea and 



China. 



Var. majus Makino. A. majus (Makino) Makino; A. clivi- 

 colum Tagawa; A. wardii var. clivicolum (Tagawa) Kurata 



O-HiROHA-NO-iNU-wARABi. Frouds larger; stipes to 50 cm. 



long; scales 10-15 mm. long, 1-1.5 mm. wide; blades 30-60 

 cm. long, rather abruptly narrowed at the tip but not long- 

 tailed; pinnae 8-10 pairs; pinnules obtuse to subacute, cuneate 

 or narrowly so at base, usually pinnately lobed on the anterior 

 side. Honshu (Kanto Distr. and westw.), Shikoku, Kyu- 

 shu. 



37. Athyrium mesosorum (Makino) Makino. Asple- 

 nium mesosorum Makino; Diplazium mesosorum (Makino) 



Koidz. NuRi-WARABi. Rhizomes creeping, to 15 cm. long, 



densely covered with the persistent stubs of old stipes, densely 

 scaly; stipes 25-40 cm. long, brown, scaly at base; scales thinly 

 membranous, pale brown, lanceolate to broadly so, 1.5-2 mm. 

 wide, filiform-acuminate, entire or sparsely fimbriate toward 

 the tip; blades herbaceous, deltoid to ovate-deltoid, 30-60 cm. 

 long and nearly as wide, short-acuminate, glabrous, subtripin- 

 nate, the rachis fulvous, lustrous; pinnae nearly opposite or 

 alternate, the upper ones narrowly oblong-ovate, 10-15 cm. 

 wide, obliquely ascending, gradually acuminate, not or slightiy 

 contracted at base, distinctiy petiolulate, equilateral except the 

 lowest one, the lowest pinnae largest; pinnules narrowly del- 

 toid-ovate, to 10 cm. long, acuminate to acute, petiolulate; ulti- 

 mate pinnules oblong to narrowly ovate, 5-25 mm. long, obmse 

 to rounded, short-petiolulate, oblique, toothed, often lobed to 

 parted, often slightly decurrent on the rachis; sori few on the 

 ultimate pinnules, nearly parallel to the costule; indusia lanceo- 

 late, 2.5-3 mm. long, entire, nearly straight, solitary, whitish. 



^Mountains; Honshu (Uzen Prov. and southw.), Shikoku, 



Kyushu. s. Korea. 



38. Athyrium crenatum (Summerf.) Rupr. var. gla- 

 brum Tagawa. Diplazium sibiricum var. glabrum (Tagawa) 



Kurata. ^Miyama-shida. Rhizomes long-creeping, 2-3 mm. 



across; stipes 20-30 cm. long, prominently scaly especially to- 

 ward the base; scales uniformly black-brown, lustrous, broadly 

 lanceolate, 7-10 mm. long, 1-3 mm. wide, sparsely spinulose 



