480 Cruciferae 



3B. Capsules dehiscent. 



5A. Capsules laterally flattened; septa narrow; valves navicular, sometimes winged on the keel. 



6A. Seeds solitary in each locule of capsule; cotyledons incumbent 4. Lepidium 



6B. Seeds 2 to several in each locule; cotyledons accumbent 5. Thlaspi 



5B. Capsules terete or dorsally flattened. 

 7A. Flowers white. 



8A. Valves elastic and coiled at maturity 6. Cardamine 



8B. Valves not elastic or coiled at maturity. 



9A. Flowers bracteate; seeds rather large, few; cotyledons incumbent 7. Wasabia 



9B. Flowers not bracteate; seeds small, many; cotyledons accumbent. 



lOA. Capsule a silique 8. Nasturtium 



lOB. Capsule an inflated silicle 9. Cochlearia 



7B. Flowers yellow. 



11 A. Cotyledons incumbent; fruit terete or flattened; valves often 3-nerved; plants hairy 10. Sisymbrium 



IIB. Cotyledons accumbent; valves costate or keeled or 1-nerved; plants glabrous. 



12A. Siliques 3-4 cm. long, costate or keeled 11. Barbarea 



12B. Siliques shorter, weakly 1-nerved or nerveless 12. Rorippa 



IB. Hairs bifid or stellate, rarely simple and gland-tipped. 



13A. Capsules flattened laterally, a silicle; valves navicular 13. Capsella 



13B. Capsules dorsally flattened or terete, a silique, rarely a silicle; valves not navicular. 



14A. Capsules a silicle; cotyledons accumbent 14. Draba 



14B. Capsule a silique. 



15A. Cotyledons accumbent; flowers white, rarely purple 15. Arabis 



15B. Cotyledons incumbent. 



16A. Flowers white or rose-purple. 

 17A. Siliques narrowly linear. 



18A. Valves thinly membranous; stamens free 16. Arabidopsis 



18B. Valves firmly membranous; longer stamens connate in pairs 17. Dontostemon 



17b. Siliques gradually narrowed above 18. Berteroella 



16B. Flowers yellow to yellowish. 



19 A. Stigma entire; leaves pinnately dissected; valves thinly membranous 19. Descurainia 



19B. Stigma bifid; leaves simple, toothed or incised; valves firmly membranous 20. Erysimum 



1. MACROPODIUM R. Br. Hakusen-nazuna Zoku 



Nearly glabrous perennial herbs; leaves alternate, toothed, the lower petiolate; inflorescence a densely flowered spikelike 

 raceme, elongate, ebracteate; flowers niany, white, rather large; sepals narrow, loose, equal; petals narrow, longer than the sepals; 

 stamens exserted, the anthers elongate, contorted; siliques elongate, flattened, long-stipitate, the valves flat, with a weak nerve; 

 style very short, the stigma punctiform; seeds in 1 or 2 series, flat, winged. Two species, in the Altai Mountains and e. Asia. 



1. Macropodium pterospermum F. Schmidt. Haku- many-flowered, often to 40 cm. long in fruit, the pedicels 



SEN-NAZUNA. Rhizomes creeping, short; stems erect, 40-100 spreading, 7-13 mm. long; petals slightly longer than the 



cm. long, usually simple, leafy, with simple minute hairs above; sepals, linear, 6-7 mm. long; stamens 13-15 mm. long; siliques 



leaves narrowly ovate to broadly lanceolate, 7-12 cm. long, 4-6 cm. long, about 4 mm. wide, the stipe 15-30 mm. long; 



1.5-5 cm. wide, acuminate, glabrous on both sides or often seeds about 10, winged on margin. July-Aug. Wet grassy 



minutely hairy on nerves beneath, toothed, the lower ones places in alpine regions; Hokkaido, Honshu (n. and centr. 



petiolate, the upper ones subsessile; inflorescence elongate, distr.) ; rare. Sakhalin. 



2. ISATIS L. Taisei Zoku 



Usually glabrous, sometimes pubescent or tomentose, often glaucescent, branched annual or biennial herbs; leaves entire or 

 obsoletely toothed, the cauline sagittate; flowers in loose ebracteate racemes, small, usually yellow; silique oblong to linear, in- 

 dehiscent, 1-seeded, flat, woody or bony on the center, coriaceous and thickened or foliaceous on margin; stigma sessile; seed 

 solitary; cotyledons incumbent. About 20 to 30 species, in Europe, N. Africa, and Asia. 



1. Isatis tinctoria L. var. yezoensis (Ohwi) Ohwi. /. wide, obtuse, shallowly toothed, sagittate; petals small, yellow; 



oblongata sensu auct. Japon., non DC; 7. japonica sensu auct. siliques cuneate-obovate, about 15 mm. long, 4-5 mm. wide, 



Japon., non Miq.; 7. yezoensis Ohwi Hama-taisei, Ezo- subacute. June-July. Seashores; Hokkaido. ^Korea and 



TAISEI. Glaucous, glabrous; stems erect, 30-40 cm. long; Ussuri. 

 lower cauline leaves narrowly ovate, 10-12 cm. long, 2-3 cm. 



3. RAPHANUS L. Daikon Zoku 



Glabrous or hispidulous branched annuals or biennials; tap-root often thickened; lower leaves lyrate; racemes elongate, bract- 

 less; flowers white or purplish rose, rather large, pedicelled; sepals erect, the lateral ones subsaccate at base; stamens free; 

 siliques elongate, terete, torulose, coriaceous or spongy, the locellus 1-seeded; style elongate, the stigma emarginate; seeds globose; 

 cotyledons conduplicate. Few species, in the Old World, especially in the Mediterranean region. 



