Cruciferae 



14. DRABA L. 



Inu-nazuna Zoku 



Usually densely tufted annuals, biennials, or perennials with stellate or simple hairs; leaves entire or incised-toothed, the radical 

 rosulate; stems naked or few-leaved; racemes short or elongate; flowers bractless, small, white or yellow, rarely rose or purple; 

 silicles elliptic to broadly linear, flat, many- to few-seeded, the valves nearly flat, rarely nerved, deciduous, the septum membran- 

 ous; styles short or elongate, the stigma simple; seeds in 2 series, wingless, sometimes with a membranous appendage; cotyledons 

 accumbent. About 270 species, mostly in mountains and northern regions of the N. Hemisphere. 



lA. Biennial; flowers yellow; style very short; silicles much shorter than the pedicels 1. D. nemorosa var. hebecarpa 



IB. Perennials; innovation-shoots present; style distinct. 



2A. Flowers yellow 2. D. japonica 



2B. Flowers white. 



3A. Petals 7-8 mm. long; style 2.5-4 mm., rarely 1.5 mm. long; silicles pilose 3. D. sachalinensis 



3B. Petals 3-6 mm. long; style less than 2 mm. long. 



4A. Silicles puberulent, prominently contorted; petals 5-6 mm. long; style 0.2-0.5 mm. long; rocks by the sea 4. D. borealis 



4B. Silicles glabrous, not or slightly contorted; petals 3-4 mm. long; rocks and dry slopes in mountains and alpine regions. 



5A. Seeds appendaged on one end S. D. sakiiraii 



5B. Seeds not appendaged. 



6A. Leaves glabrous on both sides, ciliate 6D. shiroumana 



6B. Leaves minutely stellate-pubescent on both-sides 7. D. \itadakensis 



1. Draba nemorosa L. var. hebecarpa Lindbl. D. hirta stigma capitellate; seeds not appendaged, 1-1.2 mm. long. 



sensu Miq. non L. Inu-nazuna. Rather prominendy May-June. Rocky places in mountains; Hokkaido, Honshu 



stellate-pilose biennial; stems erect, simple or branched at base, (Shinano Prov.). Sakhalin. 



10-20 cm. long, slender, leafy; radical leaves rosulate, spathu- 4. Draba borealis DC. Shirobana-no-inu-nazuna, Ezo- 

 late, 2-4 cm. long, 8-15 mm. wide, slightly toothed, nearly inu-nazuna. Rather densely stellate-puberulent perennial, 

 sessile, the cauline leaves few, narrowly ovate to narrowly the innovation shoots somewhat elongate and sparingly 

 oblong, 1-3 cm. long, 5-15 mm. wide, broadly cuneate, ses- branched; stems few-leaved, 6-20 cm. long; rosulate leaves 

 sile; racemes many-flowered, the pedicels 10-20 mm. long, spathulate, 15-30 mm. long, 5-8 mm. wide, obtuse, few- 

 spreading; petals about 3 mm. long; silicles narrowly oblong, toothed or sometimes entire, the margin with scattered sim- 

 flat, 5-8 mm. long, 2-2.2 mm. wide, short-pilose; style very pie hairs, the cauline leaves ovate to oblong, 8-25 mm. long, 

 short or nearly absent; seeds not appendaged, about 0.4 mm. toothed, usually acute, the innovation-leaves usually without 



long. Mar.-May. Sunny places in lowlands; Hokkaido, simple hairs; racemes rather many-flowered, puberulent, the 



Honshu, Shikoku, Kyushu; common. ^Temperate regions pedicels about 10 mm. long; petals obovate, clawed, 5-6 mm. 



of the N. Hemisphere. long; silicles broadly lanceolate, usually prominently con- 



Var. leiocarpa Lindbl. Me-inu-nazuna. Silicles gla- torted, 5-8 mm. long, 2.5-3 mm. wide, puberulent, rarely 



brous. Rare. Occurs widi var. hebecarpa. glabrous; style 0.2-0.4 mm. long, the stigma capitellate; seeds 



2. Draba japonica Maxim. D. yezoensis Nakai without appendage, about 1 mm. long. June-July. Rocky 



Nambu-inu-nazuna. Perennial herb, stellate-pubescent, the cliffs near the sea; Hokkaido (Nemuro and Hidaka Prov.). 



caudex branched, tufted, the innovation-shoots slightly elon- Kuriles, Sakhalin, and Ochotsk Sea region. 



gate; stems 5-10 cm. long, few-leaved; rosulate leaves oblance- 5. Draba sakuraii Makino. D. sinanensis Makino 



olate to narrowly so, 5-15 mm. long, 1.5-3 mm. wide, entire Togakushi-nazuna. Loosely mfted perennial, rather densely 



or with few coarse teeth, subobtuse, gradually narrowed below, stellate-pubescent; stems 7-20 cm. long, 2- to 5-leaved, with 



the cauline leaves sessile, 8-10 mm. long, 2-6 mm. wide; ra- scattered stellate and simple hairs, the innovation-shoots rarely 



cemes short-pilose; petals obovate, clawed below, yellow, 4-4.5 elongate; rosulate leaves oblanceolate or broadly so, 1-2.5 cm. 



mm. long; silicles narrowly ovate, glabrous, 4-6 mm. long, long, 2-8 mm. wide, toothed or subentire, subobtuse, often 



2.5-3.5 mm. wide; style 0.2-0.5 mm. long, the stigma discoid; with simple hairs on margin, the cauline leaves narrowly 



seeds not appendaged, about 1.5 mm. long. June-Aug. ovate to oblong, usually coarsely few-toothed, 8-15 mm. long, 



Alpine regions; Hokkaido (Mount Yubari), Honshu (Mount 5-10 mm. wide, puberulent, the pedicels puberulent; petals 



Hayachine); rare. 3.5-4(-5) mm. long, cuneate-obovate; silicles narrowly lanceo- 



3. Draba sachalinensis Trautv. D. grandiflora Franch., late, flat; style 1-2 mm. long, the stigma capitellate; seeds with 



non C. A. Mey.; D. jranchetii O. E. Schulz; D. moiiuana a membranous tail. June-July. Rocky dry places in high 



Koidz.; D. shinanomontana Ohwi; D. sachalinensis var. shi- mountains; Honshu (n. part of centr. distr.). 



nanomontana (Ohwi) Okuyama ^Moiwa-nazuna. Per- Var. nipponica (Makino) Takeda. D. nipponica Makino; 



ennial herb, densely grayish stellate-puberulent; stems 10-25 D. onta^ensis Makino; D. ni\oensis Koidz.; D. sa\iiraii var. 



cm. long, 2- to 4-leaved on lower half, the sterile innovation onta\ensis (Makino) Takeda; D. sa\uraii var. rigidtda Ta- 



shoots slighdy elongate, with a tuft of rosulate leaves at the keda; D. linearis Satake Kumoma-nazuna. Plant more 



tip; rosulate leaves oblanceolate, few-toothed or subentire, 2-3 slender, less puberulent, with glabrous more slender pedicels 



cm. long, 5-8 mm. wide, subacute, gradually narrowed below, and shorter-tailed seeds. Honshu (centr. distr. and Nikko). 



the cauline leaves ovate to broadly lanceolate, 1.5-2 cm. long, 6. Draba shiroumana Makino. Shirouma-nazuna. 

 3-10 mm. wide, toothed, rarely entire, cuneate; racemes rather Perennial with short slightly branched caudex; stems 5-10 

 many-flowered, stellate-puberulent, the pedicels ascending, cm. long, 1- to 4-leaved, glabrous; rosulate leaves linear-spatii- 

 15-20 mm. long; petals 7-8 mm. long; silicles broadly lanceo- ulate, 5-12 mm. long, 1.5-2 mm. wide, entire or rarely few- 

 late, stellate-puberulent, 8-10 mm. long, 2.5-3 mm. wide; toothed, glabrous on both sides, ciliate, gradually narrowed 



