Saxifragaceae 507 



8. TIARELLA L. Zuda-yakushu Zoku 



Perennial herbs with short-creeping rhizomes; radical leaves long-petiolate, simple, sometimes 3-foliolate, the cauline leaves few, 

 with small stipules; racemes or cincinni simple or sparsely branched, die bracts minute; calyx 5-lobed, subpetaloid, die calyx-tube 

 cup-shaped, adnate to the ovary at base; petals 5, small; stamens 10, filaments filiform; ovary nearly superior, l-locular, with 2 

 parietal placentae; styles slender, the stigma punctiform; capsules nearly superior, thinly chartaceous; carpels very unequal; seeds 

 few, smooth. Few species, in N. America, one in e. Asia and the Himalayas. 



1. Tiarella polyphylla D. Don. Zuda-yakushu. Rhi- branch near base, the pedicels spreading, 3-7 mm. long; flow- 



zomes creeping, branched; stolons subterranean; radical leaves ers rather numerous, white, nodding; calyx-lobes 1-2 mm. 



few, cordate-orbicular, 2-7 cm. in diameter, shallowly 5-lobed, long, narrowly ovate; petals subulate-linear, slightly longer 



obtusely toothed, long-hairy and glandular-puberulent, the than the calyx-lobes; stamens exserted; capsule-valves very 



stipules membranous, brown, the petioles 2-10 cm. long; unequal, 7-12 mm. long. June-Aug. Damp woods in 



scapes 10-40 cm. long, glandular-puberulent, with 2 or 3 mountains; Hokkaido, Honshu (Kinki Distr and northw.), 



short-petioled cauline leaves; racemes often with a short Shikoku. China, Formosa, and Himalayas. 



9. MITEULA L. Charumeru-s6 Zoku 



Low usually hirsute perennial herbs with creeping rhizomes; radical leaves long-pedolate, simple, ovate, incised to lobulate, the 

 stipules scarious; stems leafless or few-leaved; inflorescence racemose, terminal, bracteate; flowers small; calyx 5-lobed, the calyx- 

 tube cup-shaped or campanulate, more or less adnate to the ovary; petals 5, on the margin of the calyx-tube, often pinnatiparted, 

 sometimes simple or absent; stamens 5 or 10; ovary l-locular, superior or subinferior; styles 2; seeds ovoid, small, often tubercled. 

 About 20 species, in N. America, Siberia, Japan, and Formosa. 



lA. Ovary nearly superior, nearly free from the calyx-tube. 



2A. Stamens 5; petals subulate-linear, entire; stems 1- to 3-leaved 1. M. integripetala 



2B. Stamens 10; petals pinnatiparted; stems scapose or I-leaved 2. M. ntida 



IB. Ovary inferior, adnate at least partially to the calyx-tube. 



3A. Petals absent; scapes 2-7 cm. long, 1- to 3-flowered; calyx-tube campanulate 3. M. doiana 



33. Petals present; scapes 10-50 cm. long, often many-fiowercd; calyx-tube cup-shaped to broadly obconical. 



■lA. Stamens attached to the disc; leaves wider than to as wide as long; racemes few-flowered; style undivided 4. M. paucifiora 



4B. Stamens attached to the base of the petal; leaves usually longer than wide; racemes rather many-flowered; style often lobed. 



5A. Petioles and the underside of leaves glabrous 5. M. acerina 



5B. Petioles and leaves hirsute. 



6A. Calyx -lobes erect with an ascending to slightly recurved tip. 



7A. Petals entire, smooth, subulate-linear; calyx-lobes not ciliate; seeds papillose 6. M. leiopetala 



7B. Petals pinnatiparted, glandular-roughened; calyx-lobes ciliate; seeds smooth or papillose. 



8A. Leaves obtuse or subobtuse; seeds not papillose 7. M. siylosa 



8B. Leaves acute to subacute; seeds papillose 8. M. makinoi 



6B. Calyx-lobes spreading in anthesis, often reflexed at apex. 



9A. Rhizomes short, thickened, with a tuft of leaves at the apex. 



lOA. Leaves broadly ovate-orbicular, the petioles eglandular-hirsute; stipules usually glabrous. 



1 1 A. Calyx-lobes not ciliate; petals mostly 7-parted 9. M. kpshiensis 



IIB. Calyx-lobes ciliate; petals 9- to 1 1-parted 10. M. jtirusei 



lOB. Leaves usually elongate-ovate, the petioles glandular-hirsute; stipules long-ciliate 11. M. japonica 



9B. Rhizomes long-creeping, rather slender, loosely leaved. 



I2A. Seeds papillose; leaves broadly ovate; calyx-lobes deltoid; styles 2, 2-lobed 12. A/. k.'iisiana 



12B. Seeds not papillose; leaves narrowly ovate; calyx-lobes depressed -deltoid; styles 2, undivided 13. A/, yoshinagae 



1. Mitella integripetala H. Boiss. Ezo-no-charumeru- cemes few-flowered; calyx-lobes deltoid-ovate, acute, spreading, 

 so. Stoloniferous; radical leaves membranous, long-peti- subrecurved at the tip; petals 5, pinnatiparted, not glandular; 



olate, deltoid-ovate, cordate, shallowly 3- to 5-lobed, toothed, seeds glabrous, smooth, black. June. Woods in moun- 



scattered-pilose on upper side, glabrescent beneath; flowering tains; Hokkaido, Honshu (centr. distr.); rare. n. Korea, 



stems with 1-3 short-petiolate leaves; racemes rather many- Manchuria, Ussuri to Siberia, and N. America. 



flowered, the pedicels less than 3 mm. long, minutely glandu- 3. Mitella doiana Ohwi. Hime-charumeru-s6. Rhi- 



lar-puberulent; calyx-tube cup-shaped, the lobes lanceolate, zomes creeping; radical leaves tufted, ovate-cordate, 8-13 mm. 



about 2 mm. long, acute; petals linear, 4-5 mm. long, entire, long, 7-10 mm. wide, shallowly 3-lobed, acute, deeply cordate, 



minutely glandular; styles 2, subulate, enure. Hokkaido hirsute on both sides, the petioles 1-3 cm. long, hirsute, the 



(sw. distr.), Honshu (n. distr.); rare. stipules scarious, entire, glabrescent; flowering stems 2-7 cm. 



2. Mitella nuda L. Maruba-charumeru-s6. Rhizomes high, hirsute and glandular-pubescent, 1- to 3-flowered toward 

 slender, elongate; stolons filiform; radical leaves few, ovate- the top; calyx-tube subcampanulate, the lobes ovate-deltoid, 

 orbicular, 15-35 mm. long and as wide, often subcordate, ob- erect in anthesis, with a slightly recurved tip; petals absent; 

 tuse, obsoietely incised and short-toothed, scattered-pilose on styles 2, terete, about 0.8 mm. long, scarcely thickened at apex; 



both sides, the petioles 5-10 cm. long, slender, glandular- seeds papillose. May-July. Kyushu (Yakushima) ; rare. 



hairy, the stipules short; flowering stems 15-25 cm. long, 4. Mitella paucifiora Rosend. Ko-charumeru-s6. Rhi- 



short glandular-pubescent, often with a single small leaf; ra- zomes rather slender, elongate; leaves all radical, broadly 



