ROSACEAE 



537 



15. FILIPENDULA L. Shimotsuke-s6 Zoku 



Perennial herbs; radical leaves pinnate, the terminal leaflets large, palmately lobed, toothed, the lateral leaflets small or some- 

 times obsolete, the cauline leaves often without lateral leaflets, the stipules membranous or foliaceous; inflorescence a cymose 

 corymb; flowers many, small, bisexual or rarely unisexual (plants dioecious), white or pinkish; calyx-lobes 4 or 5, reflexcd; petals 

 4 or 5, orbicular; stamens rather numerous, deciduous after anthesis, the filaments filiform; carpels 3-10, rarely to 15, free, 

 usually short-stiped; ovules 2 in each carpel; style short, terminal; achenes or indehiscent follicles glabrous or ciliate, laterally 

 flattened, stiped or sessile. More dian 10 species, in temperate to cooler regions of the N. Hemisphere. 



lA. Achenes sessile, loosely contorted, semiorbicular, glabrous, rounded on outer margin at base; flowers white, unisexual; pedicels with 



crisped ascending eglandular hairs; lateral leaflets 3 or 4 pairs, very small I. F. tsiigtiwoii 



IB. Achenes stiped, not contorted, broadly oblanceolate to oblong, ciliate or glabrous, equilateral; flowers white to pink, bisexual; pedicels 

 glabrous or with spreading strict short hairs. 

 2A. Pedicels short-hairy; flowers white; achenes long-ciliate; stipules with a large semiorbicular auricle on one side. . . 2. f . \amlschauca 

 2B. Pedicels glabrous; flowers usually pink, rarely white; achenes ciliate or glabrous; stipules with or without a prominent auricle. 



3A. Stipules with a semirounded spreading foliaceous auricle on one side; lateral leaflets few-paired and very small or often obsolete. 



3. f . auricidata 

 3B. Stipules not aurided; lateral leaflets at least in the lower leaves several-paired, much smaller than the terminal leaflet but prominent. 



4. f . multijuga 



1. Filipendula tsuguwoi Ohwi. Shikoku-shimotsuke- 

 s6. Dioecious herb; stems 50-70 cm. long, erect, 5-angled, few- 

 leaved, glabrous except for short pubescence on upper part; 

 radical and lower cauline leaves long-petioled, the terminal 

 leaflets membranous, orbicular, deeply cordate, 8-13 cm. in 

 diameter, palmately 5- to 7-cleft, die lobes ovate to narrowly so, 

 long-acuminate, duplicate-toothed, green on both sides, thinly 

 pilose on nerves or glabrous on upper side, with crisped yel- 

 lowish brown pubescence near base beneath, the lateral leaflets 

 3 or 4 pairs, very unequal, the larger ones ovate, to 1-2 cm. 

 long, the smaller ones 2-3 mm. long, the sdpules of the cauline 

 leaves semi-ovate, erect, acuminate, toothed, 1-1.5 cm. long; 

 corymbs terminal and in the upper axils, the branches and 

 pedicels white-pubescent; flowers unisexual, white; calyx-lobes 

 elliptic, glabrous; petals 5, spreading, glabrous, 3.5-4 mm. 

 long; stamens about 30, usually slighriy longer than die petals 

 in the staminate flowers, much shorter than the petals in the 

 pistillate flowers; fruiting carpels 5-6, glabrous, sessile, loosely 

 contorted, nearly orbicular-ovate, about 3 mm. long, the stigma 

 small, hemispheric. July. High mountains; Shikoku, Kyu- 

 shu; rare. 



2. Filipendula katntschatica (Pall.) Maxim. Spiraea 



\amtschatica Pall. Oni-shimotsuke. Stout perennial 



herb; stems erect, 1-2 m. long, hispid or hispid-pilose, angled; 

 leaves large, die terminal leaflets orbicular, shallowly cordate, 

 15-25 cm. in diameter, palmately (3-)5-cleft, doubly incised- 

 toothed, green, hispid, the lateral leaflets few, very small, often 

 obsolete in the upper leaves, the stipules green, incised-toodied, 

 with an obliquely spreading auricle at base on one side; in- 

 florescence large, with short spreading straight hairs on pedi- 

 cels and branches; flowers white or slightly pink, 6-8 mm. 

 across; calyx-lobes reflexed; petals obovate-orbicular, nearly 

 entire; stamens 1.5 times as long as the petals; carpels 5; 



achenes oblanceolate, long-ciliate. June-Sept. Mountains; 



Hokkaido, Honshu (centr. and n. distr.) ; often gregarious 

 along streams and ravines; variable as to indument. Sometimes 

 distinguished are: Forma pilosa Koidz. vvidi looser shorter 

 hairs on stems and leaves, and forma glabra Koidz. with 

 nearly glabrous stems and leaves. 



3. Filipendula auriculata (Ohwi) Kitam. F. purpurea 



var. auriculata Ohwi. Koshiji-shimotsuke-s6. Rhi- 



zomes short and stout; stems erect, angled, 1-1.5 m. long, 

 10- to 13-leaved; radical leaves long-petioled, the terminal leaf- 

 lets large, cordate, 5-15 cm. long, 10-20 cm. wide, 5- to 7-cleft, 

 the lobes acuminate, serrulate, rarely obsoletely incised, nearly 

 glabrous on both sides or thinly pilose on nerves beneath, the 

 lateral leaflets usually 1 or 2 pairs or absent, ovate, 1-3 cm. 

 long, toothed, the stipules mosdy foliaceous, green, with semi- 

 orbicular, denticulate auricles; corymbs many-flowered, rather 

 large, glabrous; flowers pink, 4-5 mm. across; achenes broadly 



oblanceolate, ciUate, stiped. July-Aug. Low mountains; 



Honshu (Iwashiro, Echigo Prov.). 



Filipendula purpurea Maxim. Kyoganoko. Leaves 

 mostly without lateral leaflets, the stipules smaller, with a 

 smaller often indistinct auricle on one side; flowers pink; 



achenes rarely ripening. July-Aug. Long cultivated in 



gardens, and possibly a hybrid of F. multijuga and F. auricu- 

 lata. Forma albiflora (Makino) Ohwi. F. purpurea var. 



albiflora Makino Natsu-yuki-s6. Flowers white. 



Rarely cultivated in gardens. 



4. Filipendula multijuga Maxim. Spiraea palmata 

 Thunb., non L. f,, nee Pall. Shimotsoke-s6. Nearly gla- 

 brous perennial herb; stems 30-100 cm. long; terminal leaflets 

 orbicular, subcordate, 5-10 cm. in diameter, 5- to 7-cleft to 

 -parted, toothed, the lateral ones of few pairs, unequal, ovate, 

 3-30 mm. long, toothed and often incised, often reduced and 

 obscure in the upper leaves; stipules membranous, sometimes 

 few-toothed, erect or ascending; inflorescence a terminal many- 

 flowered corymb, glabrous; flowers usually rose, 4-5 mm. 

 across; calyx-lobes 4 or 5, reflexed; petals ovate-orbicular, obso- 

 letely toothed; carpels 4 or 5; achenes short-stipcd, oblong, gla- 

 brous. July-Aug. Mountains; Honshu (centr. distr. and 



westw.), Shikoku, Kyushu. 



Var. ciliata Koidz. F. ciliata (Koidz.) Miyabe & Kudo 



Akabana-shimotsuke. Achenes ciliate. Honshu 



(centr. and Kanto Distr.). 



Var. yezoensis Hara. F. yezoensis Hara Ezo-no-shi- 



M0TsuKE-s6. Calyx-lobes short-pilose inside; stipules smal- 

 ler; petals nearly entire. Hokkaido. 



16. SANGUISORBA L. Ware-moko Zoku 



Perennial herbs with stout creeping rhizomes; stems erect, leafy, ascending at base; leaves radical and cauline, alternate, 

 imparipinnate, with dilated petioles, the stipules adnate at base, the leaflets oblong or elliptic, toothed, sessile or short-petioluled; 



