Celastraceae 603 



wide, obtuse or obtusely acute, sometimes emarginate, abruptly toothed; petals orbicular, about 3 mm, long; disc cup-shaped; 



acuminate at base, entire, with slightly recurved margins, deep filaments short, borne on the margin of the disc, as long as the 



green and lustrous above, the midrib rather slender, the lateral pale yellow anthers; capsules ellipsoidal, 1.5-2 cm. long, 



nerves slender, both slightly raised, the petioles 7-12 mm. smooth, dehiscent and exposing a large red seed. Mar.-Apr. 



long; flowers 5-merous, sessile, few, on a short peduncle 7-12 Woods and thickets near seashores; Honshu (Izu, Awa, Sa- 



mm. long, pale green, about 5 mm. across, with 2 small garni Prov.), Kyushu. Ryukyus and Formosa. 



bractcoies just beneath the calyx; calyx-lobes rounded, minutely 



4. EUONYMUS L. Nishikigi Zoku 



Evergreen or deciduous, erect or scandent shrubs or trees; leaves opposite, rarely alternate, entire or serrate; flowers small, 

 bisexual or a few unisexual, in axillary cymes or solitary; calyx-lobes 4 or 5; petals attached below the disc, usually entire; 

 stamens as many as die petals and calyx-lobes, opposite the latter, the filaments sometimes absent; disc discoid, 4- or 5-lobed; 

 ovary below the disc and adnate to it, 3- to 5-locular; style short; ovules 2, rarely few in each locule; capsules 3- to 5-valved, 

 sometimes winged, rarely spinose, with 1 or 2 seeds in each locule, the seeds entirely or partially enclosed in an aril, the endo- 

 sperm fleshy. About 150 species, in temperate to warmer regions of Asia, Europe, N. and Centr. America. 



lA. Fruit divided nearly to the base into 1-4 lobes. 



2 A. Deciduous, the leaves thin, membranous I.E. alatus 



2B. Evergreen, the leaves deep green, chartaceous to thinly coriaceous 2. E. lutchuensis 



IB. Fruit entire, not lobed, sometimes winged or sulcate. 

 3A. Evergreen. 



4A. Fruits globose, not winged nor angular; branches terete. 



5A. Erect tree with smooth branches i. E. japonicus 



53. Woody climber with obsolctely punctulate branches 4. £. jortunei 



4B. Fruits angled, rather large. 



6A. Branches 4-angled 5. E. chibae 



6B. Branches terete 6. E. tanakfie 



3B. Deciduous (apparently evergreen, leaves 7-12 cm. long in E. lanceolatus) . 



7A. Filaments longer than the anthers; petioles 8-20 mm. long; peduncles short 7. E. sicboldianiis 



7B. Filaments very short; petioles 3-10 mm. long; peduncles elongate. 



8A. Branches warty; winter-buds short; inflorescence just above the leaves; fruit 4-ang!ed, narrowed toward the base. 



8. E. oligospermtis 

 8B. Branches nearly smooth; winter-buds elongate; inflorescence under the leaves; fruit rounded at base. 

 9A. Flowers 4-merous. 



lOA. Fruit with 4 broad wings; leaves obovate, 6-12 cm. long, 3-7 cm. wide; young branchlets pale green. . . 9. E. niacroplerus 

 lOB. Fruit globose, without wings or angles; leaves narrowly ovate, 6-8 cm. long, 2-3 cm. wide; young branchlets often 



whitish with waxy excrescence 10. £. yal(tishimensis 



9B. Flowers 5-merous. 



IIA. Ovary and fruit 5-locular. 



12A. Calyx-teeth denticulate; flowers purplish; fruit globose, wingless. 



13A. Stems erect; leaves ovate, thinly chartaceous, 3-8 cm. long 1 1. E. melananlhus 



13B. Stems creeping; leaves broadly oblanceolate, somewhat coriaceous, 7-12 cm. long 12. E. ianceolatus 



12B. Calyx-teeth entire; flowers white to pale purple. 



I4A. Fruit globose, wingless 13. E. oxyphyltus 



14B. Fruit with 5 short deltoid wings 14. E. planipcs 



IIB. Ovary and fruit 3- or rarely 4-locular; fruit with 3 or 4 rather broad, falcately deltoid wings near base; leaves rugulose 

 on upper side in life; flowers purple, with 5 calyx-teeth and petals 15. £. tricarpiis 



1. Euonymus alatus (Thunb.) Sieb. Celastrus alatus sublriflonis Bl.; Celastrus striatiis Thunb.; E. alatus var. sub- 



Thunb.; E. alatus var. typicus Kegel; E. striatus var. alatus triflorus (Bl.) Fr. & Sav. Ko-mayumi. Branches wing- 



(Thunb.) Nfakino Nishikigi. Horizontally spreading, less; leaves glabrous. Forma pilosus (Loes. & Rehd.) 



much-branched, deciduous shrub, the branches green, glabrous, Ohwi. E. alatus var. pilosus Loes. & Rehd.; E. thunbcrgintius 



with 4 corky wings; leaves short-petiolate, membranous, obo- Bl. Ke-nishikici. Branches winged; leaves pilose on un- 



vate to broadly oblanceolate, 1.5-7 cm. long, 1-4 cm. wide, der side. Forma apterus (Kegel) Kehd. E. alatus var. 



acuminate to acute at both ends, glabrous, finely serrulate; apterus Kegel; E. alatus var. pubcsccns Maxim., pro parte. 



cymes 1- to few-flowered, on slender peduncles about 2 cm. Ke-ko-mayumi. Branches wingless; leaves pilose on under 



long, the pedicels short; flowers 4-merous, pale green, about 6 side. 



mm. across; caly.x-teeth depressed-rounded, with minute teeth 2. Euonymus lutchuensis T. Ito. Kyukyu-mayumi. 



on margin; petals orbicular; fruiting carpels usually 1 or 2, Glabrous small evergreen tree with slender, green, 4-angled 



narrowly obovoid, about 8 mm. long, pale red, 1-seeded; seeds branchlets; leaves thinly coriaceous, broadly lanceolate, 3-7 



ellipsoidal, 4-6 mm. long, enclosed in an orange-vermilion aril. cm. long, 1-3 cm. wide, long-acuminate, obtuse to acute at 



May-June. Thickets and woods in lowlands and moun- base, remotely and obtusely serrate, the lateral nerves obsolete, 



tains; Hokkaido, Honshu, Shikoku, Kyushu; common and the petioles slender, 2-3 mm. long; inflorescence 2.5-5 cm. 



very variable. Korea, Sakhalin, s. Kuriles, Ussuri, Man- long, 1- to 3-flowered, the peduncles very slender; flowers 4- 



churia and China. Forma subtriflorus (Bl.) Ohwi. E. merous, about 6 mm. across, pale green; calyx-teeth de- 



