664 Araliaceae 



the bracts persistent; flowers greenish, 4-5 mm. across, the a slightly depressed center; seeds 4-6. Nov .-Jan. Woods 



pedicels 4-6 mm. long; calyx-teeth short; fruit subglobose, and thickets; Kyushu (s. distr. incl. Yakushima and Tanega- 



about 5 mm. across, glabrous, the disc less than 1.5 mm. across; shima). Ryukyus, Formosa, China, and Indochina. 



column of styles about 1 mm. long, the stigmas disclike, with 



5. FATSIA Decne. & Planch. Yatsude Zoku 



Evergreen shrubs without spines or prickles; leaves large, palmately 7- to 9-lobed, the petioles dilated at base; stipules absent; 

 inflorescence in large terminal panicles, the lateral umbels staminate, the terminal ones bisexual, the bracts caducous, the 

 bracteoles membranous, caducous, narrow; flowers polygamous; petals 4-6, valvate in bud; stamens as many as the petals; ovary 

 2- to 6-locular; styles of the bisexual flowers free, with a capitate stigma, of the staminate short, connate; fruit globose, fleshy, 

 the stones sulcate or rugose on back, seeds flattened. Few species, in e. Asia. 



1. Fatsia japonica (Thunb.) Decne. & Planch. Aralia white bracts in bud; umbels many-flowered, the bracteoles 



japonica Thunb.; A. sieboldii K. Koch Yatsude. Shrub, caducous, scarious, brown; flowers white, about 5 mm. across, 



unbranched or sparsely branched, with stout branches about 5-merous, the disc convex, about 3 mm. across in fruit; calyx- 



1.5 cm. thick, and large lunate leaf-scars; stems rusty-brown teeth indistinct; fruit subglobose, black, about 5 mm. across; 



woolly while young; leaves thick-coriaceous and lustrous above, styles 5, slender, about 1.5 mm. long. Oct.-Nov. Woods 



soon glabrate, nearly orbicular, 20-40 cm. across, cordate to near the sea; Honshu (Pacific side from Iwaki Prov. through 



cordate-truncate at base, long-petioled, palmately 7- to 9-lobed, Tokaido to Kii Prov.), Shikoku, Kyushu; rather common and 



the lobes narrowly ovate-elliptic, acuminate, toothed; panicles frequently cultivated, with several cultivars grown in gardens, 



terminal, 20-40 cm. long, broadly pyramidal, with caducous Ryukyus and s. Korea. 



6. DENDROPANAX Decne. & Planch. Kakure-mino Zoku 



Evergreen glabrous trees or shrubs; leaves simple, entire or 3- to 5-lobed; stipules small, intrapetiolar, adnate to the base of 

 petioles on back, sometimes absent; umbels simple or branched, the bracts small or absent, the pedicels not jointed; flowers 

 bisexual or rarely polygamous; calyx-limb distinct, entire or 5-toothed; petals rather thick, valvate in bud; stamens 5; ovary 5- 

 to 8-1 ocular; styles connate at least to middle or nearly the whole length, fruit globose or nearly so, sulcate when dried, the 



pericarp fleshy or juicy, the stones flat or subtriangular. About 30 species, in e. Asia, the Malay Peninsula, Centr. and 



S. America. 



1. Dendropanax trifidus (Thunb.) Makino. Acer trifi- plants, with a small obsolete gland beneath among the reticu- 



dum Thunb.; Hedera japonica Jungh.; Fatsia ? mitsde (Sieb.) late veinlets, the petioles 2-10 cm. long; stipules adnate to the 



de Vriese; Aralia mitsde Sieb.; Textoria japonica Miq.; D. ja- swollen base of petioles; umbels usually solitary and terminal, 



ponicus (Jungh.) Seem.; Gilibertia japonica (Jungh.) Harms; subglobose, with a flat disc 3-4 mm. across at apex of peduncle, 



G. trifida (Thunb.) Makino; T. trifida (Thunb.) Nakai the pedicels 6-15 mm. long; flowers yellowish green, 4-5 mm. 



Kakure-mino. Small glabrous evergreen tree with gray- across; calyx-teeth undulate; fruit black, broadly ellipsoidal, 



brown branches and rather thick green branchlets; leaves of in- about 8 mm. long, with a terminal disc about 2 mm. across; 



florescence broadly rhombic-ovate to ovate, 7-12 cm. long, styles 5, connate and forming a short column; seeds with 3 ob- 



3-8(-12) cm. wide, broadly cuneate at base, with 3 nerves mse ridges on back. Honshu (Kanto Distr. and westw.), 



rising from the base, entire or sometimes partly 2- or 3-lobed, Shikoku, Kyushu, 

 often deeply 3- to 5-cleft and broadly truncate at base in young 



7. ACANTHOPANAX Miq. Ukogi Zoku 



Deciduous shrubs or trees, glabrous or with crisped pubescence, the branches often prickly; leaves palmately compound, the 

 leaflets toothed; stipules often long-ciliate; umbels simple or paniculately compound; flowers polygamous or bisexual; calyx- 

 teeth minute or obsolete; petals usually 5, valvate in bud; stamens as many as the petals; ovary 2- to 5-locular; styles free or con- 

 nate to the top; fruit globose or slighdy flattened, the pericarp juicy, the stones 2-5; seeds flattened laterally. About 30 species, 



in e. and s. Asia. 



lA. Styles 2, free or connate in lower half. 

 2A. Umbels from the short spurs, with the leaves rosulate at base. 

 3A. Leaves glabrous beneath. 



4A. Leaflets herbaceous, 2-4 cm. long, rounded at apex, long-cuneate at base, the lateral nerves and veinlets prominent and slightly 



raised beneath 1. ^. japonicus 



4B. Leaflets membranous, 2-7 cm. long, acute to obtuse, acute to acuminate at base, the nerves not raised 2. A. spinosus 



3B. Leaves short-pubescent beneath, the nerves not raised 3. A. ni^aianus 



2B. Umbels terminal, on elongate loosely leaved shoots; branches with slender prickles 'i. A. trichodon 



IB. Styles 2-7, connate nearly the full length, the free portion very short. 

 5A. Styles 5-7. 



6A. Leaflets sessile, 3-7 cm. long; umbels solitary and terminal, on short spurs with rosulate leaves at base of peduncle; prickles thick; 

 styles 5- to 7-lobed at apex; flowering May and June $. A. sieboldianus 



