668 Umbelliferae 



20B. Stylopodium depressed. 

 22A. Carpels pubescent. 



23A. Carpels puberulent, widi filiform ribs 17. Seseli 



23B. Carpels pubescent, with prominent acute ribs much wider than the intervals 21. Glehnia 



22B. Carpels glabrous, with slender, acute, noncorky ribs 22. Tilingia 



19B. Carpels strongly flattened dorsally, the lateral ribs often more prominently winged than the dorsal ribs. 

 24A. Lateral ribs on carpels equaling or only very slightly broader than the dorsal ribs. 



25A. Ribs solid and subequal; calyx teeth minute or obsolete; petals at tip incurved, refuse, or bifid 24. Ligiisticum 



25B. Ribs hollow, the lateral ones membranous, winged; calyx teeth obsolete; petals only slightly inflexed and entire at apex. 



25. Coelopleurum 

 24B. Lateral ribs on carpels much broader than the dorsal ribs. 



26A. Vittae extending the full length of the carpel; petals all equal. 

 27A. Dorsal ribs on fruit winged. 



28A. Fruit-wings thick; calyx-teeth evident 26. Dystaenia 



28B. Fruit-wings thin; calyx-teeth obsolete 27. Conioselintim 



27B. Dorsal ribs on fruit scarcely winged. 



29A. Pericarp single-layered; wings on fruit thinly membranous; calyx-teeth usually evident 29. Ostericum 



29B. Pericarp multilayered; wings thicker; calyx -teeth usually minute, indistinct. 



30A. Lateral wings on fruit membranous, those of the 2 carpels not closely appressed 28. Angelica 



SOB. Lateral wings thick, those of the 2 carpels closely appressed 30. Peticedanitm 



26B. Vittae clavate, not reaching the base of the mericarp; petals unequal, the marginal ones much larger 31. Heracleum 



1. HYDROCOTYLE L. Chidomegusa Zoku 



Annual or perennial herbs with creeping or shortly erect stems usually rooting from the nodes; leaves petiolate, with mem- 

 branous stipules, herbaceous, usually simple, palmately lobed or incised, sometimes peltate; umbels capitate, small, opposite the 

 leaves or axillary, solitary or in terminal spikes, the involucre usually of small deciduous bracts, or wanting; flowers bisexual, 

 white to pale green; calyx-teeth obsolete; petals entire, acute, usually valvate in bud; disc flat; styles filiform; fruit compressed 



laterally, the commissure narrow, the carpels compressed laterally, the ribs slender, the vittae obsolete or nearly so. About 



100 species, in the Tropics and warm-temperate regions. 



lA. Leaves usually 3-6 cm. in diameter; umbels solitary to several in fascicles, sessile to long-pedunculate; fruits 15-40, 1-1.2 mm. long. 



1 . H. javanica 

 IB. Leaves 0.7—2.5 cm. in diameter; umbels solitary, pedunculate; fruits 2-10, rarely as many as 15. 



2A. Umbels cauline; stems or branches ascending to erect; leaves 2-2. 5(— 3.5) cm. in diameter; stipules prominent, brown, scarious; 

 pedicels short but evident; fruits about 1.5 mm. long. 

 3A. Leaves cordate, with margins often overlapping at base; lobes much-depressed, rounded; upper leaves often smaller and much 



shorter than the inflorescence 2. H. ramiflora 



3B. Leaves cordate with an open sinus; lobes depressed-deltoid, obtuse; upper leaves usually longer or only slightly shorter than the 



inflorescence 3. H. maritima 



2B. Umbels radical; stems creeping; leaves 5—20 mm. in diameter; stipules small and very thin; pedicels nearly lacking. 

 4A. Petioles glabrous or with a few long hairs at apex. 



5A. Leaves 0.8-1.5 cm. in diameter, often evergreen, lobed on upper j4-/4> Ac basal margins approximate; fruits few to more 



than 1 0, about 1 mm. long 4. //. sibthorpioides 



5B. Leaves 0.5—2 cm. in diameter, deciduous in winter, lobed on upper half, the basal margins not approximate; fruits few, about 



1.3 mm. long 5. H. yabei 



4B. Petioles thinly pubescent with crisped, white, recurved, short hairs; fruits about 0.6 mm. long 6. H. dichondroides 



1. Hydrocotyle javanica Thunb. H. nepalensis Hook.; leaved; leaf-blades depressed-orbicular with overlapping mar- 

 H. polycephala Wight & Arn.; H. javanica var. polycephala gins, about 2-2.5 (-3.5) cm. in diameter, the lobes much 



(Wight & Arn.) Masam. and var. laxa Masam. Oba-chi- depressed-orbicular with overlapping basal margins, rounded 



DOME. Perennial; stems long-creeping, with few-leaved as- at apex, shallowly toothed, the petioles 4-8 cm. long, the upper 



cending branches 5-25 cm. long, crisped-puberulent, sparingly ones very short, the stipules membranous, 2-2.5 mm. wide, 



branched in upper part; leaf-blades reniform-orbicular, 3-6 cm. often brownish with darker spots, toothed; peduncles 2-5 cm. 



in diameter, obtuse, deeply cordate, with callose hairs on upper long, much longer than the leaves, the pedicels short; fruits 



side, shallowly 5- to 9-lobed, the lobes depressed-deltoid, ob- more than 10, the styles slender. June-Sept. Lawns and 



tusely double-dentate, the petioles of the radical leaves 7-15 cm. sunny places in lowlands and low mountains; Hokkaido, Hon- 



long, of the cauline 5-10 mm. long, the stipules membranous, shu, Shikoku, Kyushu. Korea. 



rounded, about 3 mm. wide; umbels rather many-flowered, 3. Hydrocotyle maritima Honda. H. wilfordii sensu 



usually borne only on the branches; fruit short-pedicellate, the auct. Japon., non Maxim.; H. ramiflora var. maritima (Honda) 



styles short. July-Oct. Woods in low mountains; Honshu Hiroe No-chidome. Resembles the preceding; branches 



(Kanto Distr. and westw.), Shikoku, Kyushu. Tropics of 7-15 cm. long; leaf-blades orbicular-cordate, 2-3(-3.5) cm. in 



Asia. diameter, somewhat 5-angled, with an open sinus at base, 



2. Hydrocotyle ramiflora Maxim. H. wilfordii Maxim. palmately 5-lobed on the upper 1/2-2/3, glabrous or loosely 



0-cHiDOME. Perennial, glabrous except pubescent on long-pilose on upper side, usually long-pilose beneath, the lobes 



the upper side of leaves and petioles; stems long-creeping, slen- broadly obovate-cuneate, incised and few-toothed; peduncles 



der, with erect or ascending branches 10-15 cm. long, few- cauline, shorter (or the upper slighdy longer) than the leaves. 



