678 



Umbelliferae 



22. TILINGIA Regel Shirane-ninjin Zoku 



Glabrous perennials with thick rhizomes; stems rather slender, few-leaved, often scapose; leaves twice to several times ternately 

 compound, toothed or dissected into linear segments; umbels simple or compound, the involucral bracts 1 to few, the bracriets 

 of the involucel few small and linear; calyx-teeth deltoid, acute, small but evident; petals white, emarginate and in- 

 curved at apex; styles rather short, the stylopodium conical, entire; fruit oblong or ovoid-oblong, slightly compressed laterally, 

 tlie carpels 5-angled, suborbicular in cross section, with slender filiform ribs, the vittae solitary in the intervals, 2-4 on the 



commissure; seeds compressed, with the flat side on the commissure. Few species in e. Siberia, Manchuria, Korea, and 



Japan. 



lA. Stems 50-80 cm. long, leafy; leaves twice or once ternate, the leaflets broadly ovate to broadly lanceolate, thin, 2-8 cm. long, acuminate; 



umbels nearly glabrous, few 1 . T. holopetala 



IB. Stems 7-20 cm. long, few-leaved; leaves finely ternately dissected, the ultimate segments linear to broadly ovate, rather thick, 5-20 

 mm. long; rays minutely puberulent on the inner side. 

 2A. Ultimate leaf-segments broadly ovate to narrowly oblong, few-toothed to -cleft; styles nearly as long as or slightly longer than the 



stylopodium 2. T. ajanensis 



2B. Ultimate leaf-segments linear-filiform, entire; styles 2.5-3.5 times as long as the stylopodium 3. T. tachiroei 



1. Tilingia holopetala (Maxim.) Kitag. Carum holo- oblong lobes, 5-20 mm. long, lustrous and deep green on 

 petalum Maxim.; Ligusticum holopetalum (Maxim.) Hiroe & upper side, glabrous, the upper leaves very small, often re- 

 Constance Ibukizeri. Rhizomes ligneous, shortly duced to sheaths; umbels 1-3, the rays 5-10, 1.5-2.5 cm. long, 



branched; stems 50-80 cm. long, leafy, branched; radical and the involucral bracts 1, 2 or few, the umbellets 20- to 30-flow- 



lower cauline leaves long-petiolate, 1- or 2-ternate, the leaflets ered, the bractlets of the involucel few, linear, the pedicels 3-6 



thin, broadly ovate to broadly lanceolate, irregularly dentate mm. long, nearly as long as the involucel; flowers white, some- 



and usually pinnately incised or lobed, 2-8 cm. long, acumi- times purplish; calyx-teeth minute; styles nearly as long as or 



nate, membranous, with microscopically raised marginal cells, slighdy longer than die stylopodium; fruit ovoid, 2.5-3 mm. 



the upper leaves smaller, sessile, ultimately reduced to bracts; long; vittae 4 on the cotmnissure. July-Aug., fruiting Sept.- 



umbels few, pedunculate, nearly glabrous, the involucral bracts Oct. Alpine slopes; Hokkaido, Honshu (centr. and n. distr.). 



1 or 2, linear, the rays 8-10, 3-4 cm. long, the peduncle and Sakhalin, Kuriles, Kamchatka, Ochotsk, and e. Siberia. 



rays at apex with winglike ridges, tlie pedicels 8-12 mm. long, 3. Tilingia tachiroei (Fr. & Sav.) Kitag. Seseli tachiroei 



the involucral bractlets few, linear, 4-8 mm. long; calyx-teeth Fr. & Sav.; Cnidium tachiroei (Fr. & Sav.) Makino; Ligusti- 



evident; styles short, about as long as the stylopodium; fruit cum tachiroei (Fr. & Sav.) Hiroe & Constance ^Miyama- 



ovoid-oblong, 3.5-4 mm. long. Aug., fruiting Sept.-Oct. uikyo. Resembles No. 2; glabrous except for papillae on 



Grassy places in mountains; Hokkaido, Honshu (centr. and n. the umbels below and on the inner side of rays; rhizomes 



distr.). short, thick; stems 10-20(-35) cm. long, 1- to 4-leaved; radical 



2. Tilingia ajanensis Regel. Conioselinum \amtschati- and lower cauline leaves 3-10 cm. long, long-petiolate, deltoid- 

 cum var. alpinum Rupr.; Selinum tilingia (Regel) Maxim.; ovate, several times finely dissected into linear-filiform entire 

 Cnidium ajanense (Regel) Drude; Cnidium tilingia (Regel) segments, these 5-15 mm. long, 0.5-0.75 mm. wide, acute, en- 



Takeda; Ligusticum ajanense (Regel) Koso-Pol. Shirane- tire, smooth, the upper leaves with fewer segments or re- 



NiNjiN. Small perennial, glabrous except for papillae duced to sheaths; umbels 1-4; fruit ovoid-oblong, about 3 mm. 



on the umbels beneath and on the inner side of the rays; long; styles slender, about 1.2 mm. long, 2.5-3.5 times as long 



rhizomes short, rather thick; stems 7-20 cm. long, rarely to 35 as the stylopodium, the calyx-teeth evident; vittae 2 on the 



cm. long, leafless, or 1- to 3-leaved; radical and lower cauline commissure. Aug., fruiting Sept.-Oct. High mountains; 



leaves petiolate, 2-4 times ternately compound, the ultimate Hokkaido, Honshu (centr. and n. distr.), Shikoku (Mount 



segments broadly ovate to narrowly oblong, thick-membranous Tsurugi) . Korea. 



or herbaceous, usually with finely dissected linear to narrowly 



23. PTERYGOPLEURUM Kitag. Shimura-ninjin Zoku 



Glabrous perennial with short rhizomes and fleshy white roots; stems erect, fistulose, branched above, angular; leaves once or 

 twice ternately compound, the ultimate segments linear, entire, with thickened margins; umbels compound, the bracts and 

 bractlets few, entire, narrow; flowers white; calyx-teeth deltoid, acute; petals equal, with an incurved and shallowly bifid apex; 

 styles short, becoming recurved, the stylopodium depressed-conical with undulate margin; fruit glabrous, slighdy compressed 

 laterally, oblong, the carpels semi-orbicular in cross section, with thick, acute ribs, the vittae solitary in the intervals, 2 on the 

 cotiunissure, the carpophore bifid; seeds compressed and angular, with a flat face. One species, in Japan. 



1. Pterygopleurum neurophyllum (Maxim.) Kitag. peduncles and rays with winglike ridges at the apex, the rays 



Edosmia neurophyllum Maxim.; Carum neurophyllum 

 (Maxim.) Fr. & Sav.; Slum neurophyllum (Maxim.) Hara 



Shimura-ninjin. Stems angled, erect, loosely branched 



above, 80-120 cm. long; leaves rather firm, 2 or 3 times ter- 

 nately compound, 10-20 cm. long, deltoid-orbicular in out- 

 line, the ultimate segments linear, 4-10 cm. long, 3-5 mm. 

 wide, entire, acuminate, narrowed at base, scaberulous on the 

 margins, the upper leaves smaller and simpler, ternate, sessile, 

 directly attached by a narrow sheath; umbels compound, the 



8-10, rather firm, smooth, 2.5-4 cm. long, the involucral bracts 

 7-12 mm. long, appressed to the outer rays, bracts of the in- 

 volucel appressed to the pedicels, 2-5 mm. long, the pedicels 

 6-10 mm. long; fruit 3.5-4 mm. long, the styles about 0.5 mm. 



long, appressed to and nearly as long as the stylopodium. 



Aug.-Sept. Wet places along rivers in lowlands and plateaus; 

 Honshu (Musashi Prov.), Kyushu (Tano in Bungo); rare. 

 Korea. 



