Umbelliferae 



679 



24. LIGUSTICUM L. Maruba-toki Zoku 



Glabrous perennials; leaves ternately or pinnately compound; umbels compound, involucral bracts numerous, few, or absent, 

 bracdets of die involucel numerous, usually endre; flowers white; calyx-teeth small or obsolete; petals with an incurved 

 emarginate or bifid apex; st\'lopodium thick, sometimes conical, often undulate on the margin; fruit ovoid or oblong, nearly 

 terete to dorsally compressed in cross secdon, the commissure broad, the carpels compressed dorsally with a flat commissural 

 side, the ribs prominent, acute or winglike, the lateral ribs often broader than the dorsal, the vittae numerous, slender, some- 

 times obsolete, the carpophore bifid; seeds compressed dorsally, the seed-face flat or concave, free from the pericarp when mature. 

 About 50 species, in the N. Hemisphere. 



lA. Leaves twice ternate; rays 15-20, 2-3.5 cm. long; fruit narrowly oblong, 8-11 mm. long 1. L. hultenii 



IB. Leaves simply ternate; rays 5-8, 1-2 cm. long; fruit broadly ovate, about 2 mm. long 2. L. tsusimense 



1. Ligusticum hultenii Fern. Angelica hultenii (Fern.) 

 Hiroe; L. scothicum sensu auct. Japon., non L. Maruba- 

 toki. Rhizomes thick; stems glabrous, erect, sparsely 

 branched abo\e, terete, striate; leaves twice ternate, broadly 

 deltoid-orbicular in outline, 10-25 cm. long and as wide, the 

 ultimate segments broadly rhombic-ovate, often 3-lobed, acute 

 to obtuse, 3-5 cm. long and as wide, dentate, glabrous, the 

 upper leaves small, sessile, the uppermost reduced to short 

 sheaths; umbels few, glabrous, the peduncles with winged 

 ridges minutely serrulate in the upper portion, the rays 15-20, 

 2-3.5 cm. long, the involucral bracts few, membranous, linear, 

 8-10 mm. long, the involucel bractlets few, linear, shorter than 

 the pedicels, die pedicels 5-8 mm. long; fruit narrowly oblong, 

 8-11 mm. long, the carpels flat, semi-orbicular in cross section, 

 the ribs subequal, prominent; caly.x-teeth minute; styles short, 

 ascending, about 0.5 mm. long, the stylopodium globose- 

 conical. July-Aug., fruiting Sept.-Oct. Near the seashore; 



Hokkaido, Honshu (n. distr.). Korea, Ussuri, Sakhalin, 



Kuriles, Ochotsk, Kamchatka, and Alaska. 



2. Ligusticum tsusimense Yabe. Ostericum tsusimense 

 (Yabe) Kitag.; Pimpinella tsusimensis (Yabe) Hiroe & Con- 

 stance TSUSHIMA-NODAKE, TsUSHIMA-TOKI. RhizomCS 



short, few; roots rather thick; stems erect, branched above, 

 slender, faintly striate, 25-40 cm. long; radical and lower cau- 

 line leaves long-petiolate, deltoid or 5-angled, ternate; leaflets 

 thickish, sometimes 2- or 3-lobed, whitish beneath, coarsely 

 toothed, rhombic-ovate or the lateral obliquely ovate, 3-5 cm. 

 long, 15-25 mm. wide, acute, cuneate at base, sometimes pilose 

 above, scabrous on the margins, the veinlets slender to obso- 

 lete, the upper leaves reduced to oblanceolate sheaths; umbels 

 long-pedunculate, few, the rays 5-8, 1-2 cm. long, subequal, 

 minutely papillose on the inner side, the bracdets of die involu- 

 cel few, linear-filiform, nearly as long as the pedicels, often in- 

 curved, the pedicels about 10, 2-5 mm. long; fruit broadly 

 ovoid, about 2 mm. long, nearly round in cross section, the 



carpels with distinct, filiform, slender ribs. Aug., fruiting 



in Oct.; Kyushu (Tsushima). 



25. COELOPLEURUM Ledeb. 



EZO-NO-SHISHI-UDO ZoKU 



Tall, stout perennials; leaves large, ternately compound, the leaflets dentate; umbels compound; flowers white; calyx-teeth 

 obsolete; petals oblong, entire, incurved at apex; stylopodium flat, often undulate on the margin; fruit ovoid or ovoid-oblong, 

 slightly compressed dorsally, with a broad commissure, the carpels flat, often becoming corky on back, the dorsal ribs thick, keel- 

 like, the lateral winged, the vittae numerous, the carpophore bifid; seeds flattened dorsally, nearly plane on face. Few species, 



in Siberia and w. N. America. 



lA. Stems often more than 1 m. long; rays numerous, densely pubescent; pedicels 4-7 mm. long; fruit often pubescent while young. 



1 . C. lucidum 

 IB. Stems usually 40-70 cm. long; rays 20-30, densely puberulcnt; pedicels 1-1.5 cm. long, slender, smooth; fruit glabrous. 



2. C. muUisectum 



1. Coelopleurum lucidum (L.) Fern. var. gmelinii 

 (DC.) Hara. Archangelica gmelinii DC; Angelica gmelinii 



(DC.) Wormsk. ex DC. in syn.; C. gmelinii (DC.) Ledeb. 



Ezo-No-sHisHi-UDO. Stout, 1-1.5 m. high, glabrous except short- 

 pubescent on the upper part; leaves twice ternate, the ultimate 

 segments ovate to rhombic-ovate, usually cuneate at base, often 

 3-lobed, mostly regularly dentate, pilose on the ncr\'es, the up- 

 per leaves reduced to inflated sheaths; involucral bract soHtary 

 or absent, densely short yellowish brown pubescent on upper 

 part, the rays numerous, densely pubescent, unequal, the in- 

 volucel-bractlets many, broadly linear, nearly as long as the 

 pedicels, the pedicels 4-7 mm. long; calyx-teeth absent; fruit 

 oblong, 6-8 mm. long, about as long as or longer than the 

 pedicels, often pubescent when young, with nearly equal, 

 thick, corky ribs, the lateral ribs slighdy broader than the 

 dorsal, the vittae solitary in the intervals, the ribs 2 on the 



commissure; seed drupelike, with a slightly convex face. 



Hokkaido.- — —Kuriles, Sakhalin, Ochotsk, and Kamchatka. 



Var. trichocarpum (Hara) Hara. Angelica trichocarpa 

 Hara; C. trichocarpum (Hara) Kitag. Ezo-yama-zenko. 



Involucre of several bracts; leaflets smaller, 1.5-3.5 cm. long, 



irregularly acutely dentate. Aug. Mountains; Hokkaido. 



2. Coelopleurum multisectum (Maxim.) Kitag. An- 

 gelica muhisecta Maxim. Miyama-zenko. Roots thick; 



stems erect, 40-70 cm. long, sparsely branched, stout, striate, 

 glabrous, except pubescent below the umbels; leaves 3 to 5 

 times ternate, deltoid in outline, 10-15 cm. long, glabrous, the 

 leaflets rather thin, oblong, ovate or rhombic-ovate, 1-3 cm. 

 long, 7-20 mm. wide, acuminate, irregularly dentate, the upper 

 leaves small, reduced to inflated sheaths; umbels few, long- 

 pedunculate, the rays 20-30, spreading, densely puberulent, 

 4-6 cm. long, bractlets of the involucel as long as to shorter 

 than the pedicels, the pedicels 20-40, smooth, slender, 1-1.5 

 cm. long; fruit oblong, glabrous, somewhat compressed dor- 

 sally, 4.5-5 mm. long, the ribs prominent, rather corky, thick- 

 ened, the lateral ones slightly broader than the dorsal, the 

 vittae solitary in the intervals, 2-^ on die commissure, the 

 sr>'les slender, about 1.5 mm. long, the stylopodium conical, 



the calyx-tccth obsolete, the carpophore bifid. July-Aug., 



fruiting in Sept. Alpine; Honshu (centr. distr.). 



