680 Umbelliferae 



26. DYSTAENIA Kitag. Seri-modoki Zoku 



Stems rather stout; radical and cauline leaves petiolate, 2 or 3 times ternately pinnate, the ultimate segments toothed and in- 

 cised, often 3-lobed; umbels compound, the involucral bracts lacking, the involucel of many, narrow bracdets; flowers white; 

 calyx-teeth evident, acute; petals equal, deeply emarginate at the incurved apex; stylopodium depressed-conical, undulate on 

 the margin, the styles slender; fruit broadly ellipsoidal, compressed dorsally, with a broad commissure, the dorsal ribs thin, 

 winged, the lateral more broadly winged, the vittae irregular, weak, numerous between and beneath the ribs, seeds semi- 

 orbicular, with a flat face, the carpophore bipartite. ^Two species, in Japan and Korea. 



1. Dystaenia ibukiensis (Yabe) Kitag. Ligusticum long, 1-2 cm. wide, pinnately lobed or incised and dentate, 



ibu\iense Yabe; Angelica ibu\iensis Makino ex Yabe Seri- acuminate, scaberulous on the nerves and margins, the veinlets 



MODOKi, Tani-seri-modoki. Perennial herb, white-papillose prominent; umbels few, the rays 10-20, 2-4 cm. long, the 

 beneath the stem-nodes and in the inflorescence; stems 30-70 involucel of filiform-linear, scabrous bractlets slightly longer 

 cm. long, striate, branched above; leaves twice ternately pin- than the pedicels, the pedicels 5-10 mm. long; fruit 5-6 mm. 

 nate, deltoid in outline, 10-25 cm. long, the ultimate segments long, the styles reflexed, 1.5-2 mm. long. July-Aug., fruit- 

 narrowly rhombic-ovate to deltoid-ovate or orbicular, 2-4 cm. ing Sept.-Oct.; Honshu (centr. and n. distr.). 



27. CONIOSELINUM Fisch. Miyama-senkyu Zoku 



Perennials with fistulose stems; leaves twice ternately pinnate, the ultimate segments pinnatifid; umbels compound, the rays 

 numerous, the involucral bracts few or absent, the bractlets of the involucel narrow; flowers white; calyx-teeth usually obsolete; 

 petals obovate with an incurved and emarginate apex; stylopodium conical; fruit oblong, compressed dorsally, the carpels flat, the 

 ribs membranous, winglike, the lateral much broader than the dorsal, the vittae 1 to few in the intervals, 4-8 on the com- 

 missure, the carpophore bifid; seed with a plane face. Species few, in the N. Hemisphere. 



lA. Primary leaflets acute to short-acuminate, firmly membranous; fruit oblong, the dorsal wings broad 1. C. kamtschaticum 



IB. Primary leaflets long-acuminate, thinly membranous; fruit ellipsoidal with narrowly winged dorsal ribs 2. C. fiUcinum 



1. Conioselinum kamtschaticum Rupr. C. gmelinii 2. Conioselinum filicinum (H. Wolff) Hara. C. uni- 



sensu Hult., non Coult. & Rose ^Karafuto-ninjin. Peren- vittatum sensu auct. Japon., non Turcz.; C. nipponicum Hara; 



nial, glabrous except the inflorescence; stems striate, simple or Peucedanum filicinum H. Wolff Miyama-senkyu. Rhi- 



sparingly branched above; rhizomes short, thick, with thick- zomes stout; stems striate, glabrous except the inflorescence, 



ened roots; radical and lower cauline leaves long-petiolate, sparsely branched above, 40-80 cm. long; radical and lower 



triternate, deltoid in ouriine, 10-20 cm. long, the primary cauline leaves thinly membranous, long-petiolate, ternately tri- 



leaflets acute to short-acuminate, the ultimate segments nar- pinnate, deltoid in outiine, 15-25 cm. long, the primary leaflets 



rowly to broadly ovate, pinnatifid, acute, incised-dentate, 15- long-acuminate, the ultimate segments narrowly ovate or ob- 



35 mm. long, 1-2 cm. wide, scaberulous on the margins, the long, acute to acuminate, pinnatifid, 2-5 cm. long, 1-3 cm. 



veinlets distinct, the upper leaves small, reduced to basal wide, incised-dentate, scaberulous on the margins, the veinlets 



sheaths; umbels few, the involucral bracts few, linear, spread- slender; umbels few, the rays 15-25, 4-6 cm. long, papillose- 



ing, shorter than the rays or lacking, the rays 10-20, 2-3 cm. puberulent on the inner side and on the upper part of the 



long, papillose on the inner side and on the upper part of the peduncle, the involucral bracts 2 or 3, linear, 1-2 cm. long or 



peduncle, the involucel of a few, linear bractlets as long as or lacking, the bractlets of the involucel few, filiform-linear, as 



slightiy shorter than the flowers, the pedicels 20-30, 6-8 mm. long as to slightly longer than the flowers, the pedicels 7-10 



long; fruit oblong, about 4 mm. long, the dorsal wings broad, mm. long, the styles about 1 mm. long, slender, recurved; 



the styles slender, appressed to the stylopodium, about 1 mm. fruit elliptic, 4-5 mm. long, the dorsal wings narrow; vittae 



long; vittae 2 or 3 in the intervals, 6 on the commissure. solitary in the intervals, 2 on the commissure. Aug., fruit- 



Aug., fruiting in Oct. Grassy places near the sea; Hokkaido. ing in Oct. Alpine regions; Hokkaido, Honshu (centr. and n. 

 Sakhalin, Kuriles, Kamchatka and e. Siberia to Alaska. distr.) . s. Kuriles. 



28. ANGELICA L. Shishi-udo Zoku 



Usually stout perennial herbs with robust, leafy, fistulose or rarely solid stems; leaves ternately or pinnate-ternate compound, 

 the ultimate segments often large, dentate; umbels compound, few to numerous, with numerous rays, the bracts and bractlets 

 small or lacking; flowers white to dark purple; calyx-teeth minute or obsolete; petals obovate to oblanceolate, narrowed at base, 

 usually with an incurved and emarginate apex; stylopodium flat, entire; fruit ovate, dorsally much compressed, with a broad 

 commissure, the carpels lunate or planoconvex in cross section, the lateral ribs membranous, winged, the dorsal usually prom- 

 inent but unwinged, the vittae 1-3 in the intervals, 2 to several on the commissure, the carpophore bifid; seeds compressed with 

 a flat or slightly concave face. More than 50 species, in temperate regions of the N. Hemisphere. 



lA. Leaves simply pinnate; leaflets lanceolate with regular minute teeth, the lower often 2- or 3-lobed; sheaths of upper leaves somewhat 



inflated; carpels semiorbicular in cross section, with very narrowly winged lateral ribs \. A. cartilaginomarginata 



IB. Leaves usually ternately or pinnate-ternate compound; carpels strongly compressed dorsally, planoconvex in cross section, with promi- 

 nently winged lateral ribs. 

 2A. Small or medium-sized herbs; leaf -segments 0.5-7 cm., rarely to 10 cm. long. 

 3A. Leaflets thick or thickish, not herbaceous. 



4A. Ultimate leaf -segments dissected into linear to lanceolate lobes 2-5 mm. wide; upper leaves reduced to much-inflated sheaths. 



