814 Lentibulariaceae 



branched, erect, rarely elongate and scandent, often scaly, 1- to many-flowered; flowers on a terminal raceme, the bracts small, 

 the pedicels solitary, often 2-bracteolate; calyx 2-toothed or -parted, often slightly accrescent after anthesis; corolla often incurved, 

 spurred, the upper lip erect, entire or 2-lobed, the lower lip usually larger, spreading, 2- to 3-lobed or again lobulate; filaments 

 often thickened, incurved; stigma-lobes lamellate, sometimes reduced to a single lobe; capsules irregularly dehiscent or 2-valved; 



seeds ovoid, compressed above or lenricular, reticulate, striate, or rugulose, sometimes spinulose. About 250 species, in wet 



muddy soil, or in ponds and lakes. CosmopoUtan. 



lA. Plants terrestrial, of wet muddy places; aerial leaves linear or spathulate, simple. 



2A. Bracts and scalelike leaves on scapes oblanceolate, dorsilixed; calyx minutely mammlllate externally; spur incurved; plant becoming 



blackish when dry I- U. racemosa 



2B. Bracts and scalelike leaves on scapes lanceolate or ovate-elliptic, basifixed; calyx smooth; spur outwardly curved; plants not be- 

 coming blackish when dry. 

 3A. Scapes 1-3 cm. long; flowers 2-3 mm. across, nearly sessile; capsules about 1 mm. long, nearly as long as the calyx. 



2. TJ. nipponica 

 3B. Scapes 5-20 cm. long; flowers 4-5 mm. across, pedicelled; capsules 3-4 mm. long, shorter than the calyx. 



4A. Flowers pale purple, rarely white; leaves spathulate 3. U. yakitsimensis 



4B. Flowers yellow; leaves linear 4. U. bifida 



IB. Plants aquatic; leaves all submerged, finely dissected; aerial leaves absent. 



5A. Leaves of two kinds, the floating ones trichotomously or dichotomously dissected, 5-10(-15) mm. long, the rachis not elongate, 

 the immersed leaves less dissected, bladder-bearing; pedicels not reflexed in fruit; plants growing in mud. 

 6A. Floating leaves capillary, with 3-5 segments, nearly bladeless in the immersed ones, each leaf uniformly with 1-3 bladders. 



5. U. exoleta 

 6B. Floating leaves dissected into 5-10, flat, linear lobes without bladders, the immersed ones with reduced blades and with well- 

 developed bladders. 



7A. Floating leaves with rather narrow entire segments 6- U. multispinosa 



7B. Floating leaves with rather broad segments loosely and minutely toothed on upper half I.U. intermedia 



5B. Leaves all alike, 2-5 cm. long, with an elongate rachis, pinnately much-dissected into narrow segments; plants free floating. 

 8A. Leaves always with well-developed bladders; scapes without cleistogamous flowers at base. 



9A. Pedicels 6-10(-12) mm. long, deflexed and thickened above after anthesis; scapes without scalelike leaves; spur and upper 



lip of corolla minutely puberulent 8. f7. pilosa 



9B. Pedicels 1.5-2.5 cm. long, arcuate after anthesis, not thickened; scapes usually with few scalelike leaves; corolla glabrous. 

 lOA. Floating stems thicker than the scape, with slender leafless respiratory branches; flowers about 15 mm. across, sterile. 



9. v. japonica 

 lOB. Floating stems more slender than the scape, whhout respiratory branches; flowers about 10 mm. across, fertile; capsules 

 globose. 

 11 A. Spur longer than the lower lip, about 8 mm. long; teeth and spinules of leaf -segments often obscure. 



10. U. sia\ujiiensis 



IIB. Spur shorter than the lower lip; teeth and spinules of leaf-segments distinct 11. U. tenuicaulis 



8B. Leaves usually bladderless, occasionally with small bladders; scapes with solitary single cleistogamic flowers at base and on 

 adjacent nodes of floating stems 12. C/. dimorphantha 



1. Utricularia racemosa Wall, ex Walp. U. caemlea long as to slightly longer than the bracts; calyx smooth, about 



Wight, non L.; Calpidisca ta\ena\ae Nakai Hozaki-no- 1 mm. long, enclosing the capsule in fruit, the lobes not ac- 



MiMiKAKiGUSA. Delicate herb of wet muddy places, di- crescent after anthesis, rounded and nearly all alike; corolla 



morphic; subterranean stems capillary, with 1 to few remote purplish, 2-3 mm. across, the spur curved forward, longer 



aerial leaves, these simple, minute, narrowly oblanceolate to than the lower lip, obtuse; capsules broadly ellipsoidal, about 



narrowly obovate-spathulate; scapes slender, 10-30 cm. long; 1 mm. long, as long as the calyx; seeds minute, smooth, ovoid, 



scales few, loose, appressed to the scape, oblanceolate, 2-3.5 wingless. Aug.-Oct. Wet muddy places in lowlands; Hon- 



mm. long, long-acuminate at both ends, dorsifixed; flowers shu (Tokaido and Ise Prov.) ; rare. 



4-10, rather loose, the bracts simulating the scales, the bracte- 3. Utricularia yakusimensis Masam. V. yaktisimensis 



oles linear, 1-1.5 mm. long, as long as the pedicels; calyx f. albida (Makino) Hara; U. affinis sensu auct. Japon., non 



broadly ellipsoidal, about 2.5 mm. long, densely mammillate Wight— — Murasaki-mimikakigusa. Glabrous slender herb; 



externally; corolla purple, about 4 mm. across, the spur about subterranean stems capillary, with bladders; scapes erect, very 



twice as long as the lower lip, incurved; capsules globose, 2.5-3 slender, 7-15 cm. long; aerial leaves small, oblanceolate or 



mm. across, enclosed in and as long as the persistent calyx; narrowly so, very obtuse; scalelike leaves few, ovate to nar- 



style about 0.5 mm. long; seeds wingless. June-Sept. Wet rowly so, membranous, obtuse or acute, 0.5-0.8 mm. long, 



muddy places in lowlands and hills; Hokkaido, Honshu, Shi- basifixed; flowers 1-3 (^), loose, the bracts similar to die scale- 



koku, Kyushu; rather common. Korea, China, Formosa, like leaves, slightly longer, 1-1.5 mm. long, acute, the bracte- 



and India. oles 2, linear, shorter than tlie bracts, the pedicels erect or sub- 



2. Utricularia nipponica Makino. Hime-mimikaki- erect, 2-3 mm. long in anthesis, 3-6 mm. long in fruit, winged; 

 GUSA. Subterranean stems capillary, loosely bladder-bearing; calyx-segments membranous, broadly ovate, obtuse, 2-3 mm. 

 aerial leaves distant, linear, obtuse; scapes 1-3 cm. long, very long at anthesis, accrescent and 4-5 mm. long in fruit, erect, 

 delicate, erect, 1- to 3-flowered; scalelike leaves 2 or 3, loose, entire, enclosing the capsule; corolla 3.5-4 mm. across, pale 

 narrowly deltoid, basifixed, acuminate, about 0.5 mm. long, the purple, rarely white, the spur 2-3 mm. long, directed down- 

 bracts basifixed, rather membranous, small, the pedicels as ward, obtuse; capsules broadly ellipsoidal or globose, about 3 



