COMPOSITAE 



859 



4. LEONTOPODIUM R. Br. 



UsU-YUKI-SO ZOKU 



Low tufted white-woolly perennial herbs, rarely woody at the base; leaves entire, alternate, lingulatc to linear; heads hetero- 

 gamous, often unisexual, in a dense hcadlLke corymb surrounded by many spreading bracteate leaves; involucres subglobose or 

 campanulate-globose, the bracts 3-seriate, nearly equal, erect, oblong, scarious and brown on margin, usually woolly on back; 

 pistillate florets marginal, filiform, the corollas broadened at tip, 5-toothed; staminate florets central, tubular, the corollas 

 slightly broadened at tip; styles entire; anthers sagittate, caudate; achenes oblong, flattened; pappus-bristles slender, 1-seriate, 

 short-setulose or scabrous, slightly connate at base. ^More than 30 species, in Eurasia and S. America. 



lA. Caulinc leaves lanceolate to narrowly oblong; stems rather many-leaved 1. L. japonicum 



IB. Cauline leaves linear to narrowly oblanccolate; stems few-leaved. 

 2A. Cauline leaves or some of them sheathing at base. 



3A. Leaves of innovation shoots linear-lanceolate, 3-8 cm. long 2. L. hayachinense 



3B. Leaves of innovation shoots linear to oblanceolate. 



4A. Flowering stems 6—15 cm. long; heads 4-10; achenes about \.2 mm. long, pilose 3. L. jauriei 



4B. Flowering stems 4-7 cm. long; heads 2-3; achenes about 1.5 mm. long, glabrous 4. L. shinanense 



2B. Cauline leaves long-petiolate, not sheathing at base 5. L. discolor 



1. Leontopodium japonicum Miq. Gnaphdlum sie- 



boldianum Fr. & Sav. Usu-yuki-so. Rhizomes tufted; 



stems 25-55 cm. long, leafy to the top; median cauline leaves 

 lanceolate to narrowly oblong, 4-6.5 cm. long, 5-15 mm. wide, 

 acute to acuminate, abruptly narrowed at base, sessile, green 

 and glabrous or thinly woolly above, whitish woolly beneath; 

 bracteal leaves loosely arranged, smaller than the upper leaves, 

 sordid-yellow tomentose on upper surface; involucres 4—5 mm. 

 long, about 5 mm. across; bracts acute or acuminate; achenes 



about 1 mm. long, more or less papillose. July-Oct. 



Hokkaido, Honshu. China. Forma orogenes (Hand.- 



Mazz.) Ohwi. L. japonicum var. orogenes Hand.-Mazz. 



Yama-usu-yuki-so. Stems 20-40 cm. long; leaves about 3 

 cm. long, 6-9 mm. wide; inflorescence branched; heads many. 



Kyushu, Honshu. Forma pemiveum (Honda) Ohwi. 



L. pemiveum Honda Kawara-usu-yuki-s6. Leaves about 



2 cm. long, densely tomentose. Mountains; Honshu (Shi- 



nano Prov.). 



Var. shiroumense Nakai ex Kitam. L. japonicum forma 



shiroumense (Nakai) Ohwi Takane-usu-yuki-so. Plants 



to 10 cm. high, with smaller leaves and fewer heads. Al- 

 pine regions; Honshu. 



Var. spathulatum (Kitam.) Murata. L. spathulatum 



Kitam.; L. japonicum forma spathulatum (Kitam.) Ohwi 



Ko-usu-YUKi-s6. Plants quite low, tufted and with smaller 



spathulate leaves. Alpine regions; Honshu (Yamato Prov.) 



and Shikoku. 



2. Leontopodium hayachinense (Takeda) Hara & 

 Kitam. L. alpinum subsp. campestre var. hayachinense Ta- 

 keda; L. discolor var. hayachinense (Takeda) Takeda & 



Beauverd Hayachine-usu-yuki-s6. Stems 10-20 cm. long, 



loosely leaved; leaves of innovation shoots linear-lanceolate, 

 3-8 cm. long, 3-5 mm. wide, usually acute, green and woolly 

 on upper surface, gray-woolly beneath; radical leaves of flow- 

 ering stems and lower cauline leaves small, the median leaves 

 erect, lanceolate, 3-5 cm. long, 4-6 mm. wide; bracteal leaves 

 5-15, linear-lanceolate to narrowly oblong, 7-30 mm. long, 1-6 

 mm. wide, acute to acuminate; heads 4-8; involucres globose, 

 about 5 mm. long; bracts obtuse, woolly; achenes about 1.6 



mm. long, papillose. July-Aug. Alpine regions; Honshu 



(Mount Hayachine in Rikuchu). 



3. Leontopodium fauriei (Beauverd) Hand.-Mazz. L. 

 alpinum sensu auct. Japon., pro parte, non Cass.; L. alpinum 

 subsp. fauriei Beauverd; L. alpinum var. fauriei (Beauverd) 



Beauverd Miyama-usu-yuki-so, Hina-usu-yuki-so. Flow- 

 ering stems 6-15 cm. long, woolly; leaves of innovation shoots 

 linear-oblanceolate, 2.5-6 cm. long, acute, long-petiolate, slightly 

 woolly on upper surface, densely so beneath; lower cauline 

 leaves linear, 1.5-3 cm. long, 2-3 mm. wide, gradually nar- 

 rowed at base, sheathing the stems, densely yellowish gray 

 woolly; bracteal leaves 8-13, linear, to 22 mm. long, 2{-A) 

 mm. wide; heads densely arranged, 4-10; involucres globose, 

 about 4 mm. long; bracts acute, long-hairy on back, with 

 brown scarious margins; achenes about 1.2 mm. long, 4-angled, 



pilose. July-Aug. Alpine regions; Honshu (n. distr.). 



Var. angustifolium Hara & Kitam. Hosoba-hana-usu- 

 YUKi-so. Plant more slender, with narrower leaves 1-1.5 



mm. wide. Honshu (Mount Shibutsu and Tanigawa in n. 



Kanto Distr.). 



4. Leontopodium shinanense Kitam. L. alpinum 



sensu auct. Japon., non Cass. Hime-osu-yuki-s6, Koma-usu- 



YUKi-so. Flowering stems slender, 4-7 cm. long, woolly; 

 leaves of irmovation shoots oblanceolate, 8-20 mm. long, 2-4 

 mm. wide, usually acute, long-petiolate, loosely gray-woolly on 

 upper surface, densely so beneath; radical and lower cauline 

 leaves spathulate or oblanceolate, the median narrowly lin- 

 gulatc, 14-20 mm. long, 2.5-3 mm. wide, obtuse, sheathing the 

 stems, densely gray-woolly on both sides; bracteal leaves 6-9, 

 lanceolate, 7-14 mm. long, 1-3.5 mm. wide, acute; heads 2 or 

 3; involucres about 4 mm. long; bracts acute or acuminate, 

 with brownish scarious margins; achenes about 1.5 mm. long, 



glabrous. July-Aug. Alpine regions; Honshu (Mount 



Nishi-koma in Shinano). 



5. Leontopodium discolor Beauverd. L. japonicum 

 subsp. sachalinense Takeda; L. sachalinense (Takeda) Miyabe 



& Kudo Ezo-usu-YUKi-s6. Flowering stems 1.3-33 cm. 



long, woolly; leaves of the innovation shoots linear-lingulate, 

 3.5-8 cm. long, 5-7 mm. wide, acute, mucronate, long-petio- 

 late, glabrous or thinly woolly and green above, whitish 

 woolly beneath; median cauline leaves lanccolatc-lingulate, 

 .3-5.5 mm. long, (1.5-)5-8 mm. wide, long-narrowed and 

 petiolelike at base, glabrous to loosely white-woolly on upper 

 surface, densely whitish woolly beneath; bracteal leaves lance- 

 olate to narrowly oblong, acute to acuminate at both ends; 

 heads 5-22, densely arranged; involucres 3-4 mm. long; bracts 

 acute to obtuse, brown, scarious on the margins, densely 

 woolly on back; achenes about 1 mm. long, 4-angled, glabrous. 

 July-Sept. Hokkaido. Sakhalin. 



