864 



COMPOSITAE 



lA. Stems woody and branched; rhizomes indistinct; heads solitary and terminal on the branchlets. 



2A. Leaves ovate, appressed-pilose; heads on elongate branchlets 1. P. scandens 



2B. Leaves oblong, nearly glabrous; heads on abbreviated branchlets 2. P. glabrescens 



IB. Stems herbaceous, simple; rhizomes elongate; heads in racemes, spikes, or panicles. 

 3A. Leaves long-petiolate. 



4A. Leaves 3-lobed; heads 1-flowered, in panicles; achenes short-pilose; receptacle glabrous 3. P. triloba 



4B. Leaves dentate or irregularly incised; heads few-flowered, in spikes; achenes glabrous; receptacle setose 4. P. robusta 



3B. Leaves sessile, obovate-oblong, 10-30 cm. long, cuneate at base 5. P. rigidiila 



1. Pertya scandens (Thunb.) Schultz-Bip. Erigeron the petioles to 12 cm. long; leaves of the inflorescence sessile, 



scandens Thunb.; P. ovata Maxim. Koya-boki. Stems small, ovate; heads in spikes, few-flowered, sometimes gemi- 



slender, woody, short-pilose, gray-brown, much branched; nate; receptacles setose, the hairs 0.5-1 mm. long; involucres 



leaves on previous season's branches many, loosely arranged, 17-27 mm. long, 8-18 mm. across; bracts many-seriate, rather 



ovate, 2-5 cm. long, 1.5-4.5 cm. wide, obtuse to acute, rounded coriaecous, the outer depressed-ovate, rounded at apex, the 



at base, subsessile, loosely mucronate-toothed, 3-nerved, ap- median ovate, purplish brown, the inner narrowly oblong; 



pressed-pilose; leaves of current season 3 or 4, fasciculate, de- achenes 10-11 mm. long, glabrous; pappus 13-15 mm. long, 



ciduous; heads solitary, terminal; involucres 13-14 mm. long; the bristles minutely scabrous, unequal. Sept.-Nov. 



bracts distincdy imbricate, acute, the outer ovate, the median Woods in mountains; Honshu, Shikoku, Kyushu, 



ovate-oblong, the inner narrowly oblong; florets about 13, Var. kiushiana Kitam. Macroclinidium robustum var. 



about 15 mm. long; achenes densely pilose, about 5.5 mm. \iushianum (Kitam.) Honda Tsukushi-kashiwaba-ha- 



long. Sept.-Nov. Thin woods in hills and low moun- guma. Heads short-pedunculate; leaves often broader; in- 



tains; Honshu, Shikoku, Kyushu. China. volucral bracts furfuraceous-puberulent on back. Kyushu. 



2. Pertya glabrescens Schultz-Bip. P. scandens sensu 5. Pertya rigidula (Miq.) Makino. Eupatorium rigid- 

 auct. Japon., non Schultz-Bip.; P. glabrescens var. viridis Na- ulum Miq.; Macroclinidium rigidulum (Miq.) Makino; M. 



kai; P. scandens var. schultziana Franch. ^Nagaba-koya- verticillatum Fr. & Sav. Kukumaba-haguma. Rhizomes 



BOKi. Rhizomes thickened; stems slender, to 120 cm. long, creeping, knotty; stems 60-90 cm. long, nearly glabrous; 



nearly glabrous, much branched; leaves on previous season's leaves coriaceous, mostly 7 or 8 in false verticils near the 



branches ovate, 25-35 mm. long, 15-25 mm. wide, obtuse to median portion of the stems, obovate-oblong, 10-30 cm. long, 



acute, rounded at base, very short-petiolate, mucronate-toothed, 3.5-11.5 cm. wide, mucronate, cuneate at base, mucronate- 



glabrous on both sides, 3-nerved; leaves of current season 3-5, toothed, subtrinerved, sessile; leaves of the inflorescence small, 



fasciculate on the nodes of the last year's branches, oblong, broadly lanceolate; heads on slender peduncles, in panicles, 



3-6.5 cm. long, 1-1.7 cm. wide, acute at both ends, serrulate, bracteate; florets 7-9; involucres 18-20 mm. long, about 1 cm. 



lustrous, sparsely pilose; heads terminal on short spurs; flor- across; bracts rounded at apex, the outer 1.5-2 mm. long, 



ets 15-18 mm. long; achenes 6.5-7 mm. long, 10-ribbed, ap- depressed-ovate; achenes 6-8 mm. long, coarsely pilose. 



pressed-pubescent. Aug.-Nov. Thin woods in hUls and Oct.-Nov. Woods in mountains; Honshu (Kinki Distr. and 



low mountains; Honshu, Shikoku, Kyushu. eastw.) . 



3. Pertya triloba (Makino) Makino. Macroclinidium The following interspecific hybrids are reported: 



trilobum Makino; P. jauriei Franch. Oyari-haguma. Pertya X hybrida Makino. Macroclinidium hybridum 



Stems erect, 45-85 cm. long, nearly glabrous; radical and (Makino) Matsum.; P. macrophylla Nakai; Macropertya hy- 



lower cauline leaves scalelike; median leaves alternate, charta- brida (Makino) Honda Kakoma-haguma, Oba-koya-boki. 



ceous, oblong, 10-13 cm. long, 7-13 cm. wide, mucronate, 3- Annual; median leaves about 8 cm. long, 6 cm. wide, 3-nerved, 

 lobed, long-petiolate, coarsely mucronate-toothed, glabrescent the petioles about 1 cm. long; upper leaves small; heads soli- 

 above, crisped-puberulent on nerves and veinlets beneath and tary in axils of upper leaves, forming a terminal spike; involu- 



on margin; upper leaves gradually smaller, becoming bracdike; cres 17-19 mm. long. Hybrid of P. robusta X P. scandens. 



heads many, paniculate, l-flowered, subsessile; receptacles gla- Pertya X suzukii Kitam. Macroclinidium suzukiiYAxs.m.., 



brous; involucres 14-17 mm. long, narrowly tubular; bracts in syn. Iwaki-haguma. Median leaves chartaceous, ap- 



ovate, the median oblong, the inner linear-oblong; achenes proximate, irregularly dentate; florets 3 or 4. Honshu 



8.5-9 mm. long, short-pilose. Sept.-Oct. Woods; Honshu (Iwaki Prov.). Hybrid of P. triloba X P. rigidula. 



(n. distr. and n.Kanto Distr.). Pertya X koribana (Nakai) Makino & Nemoto. Macro- 



4. Pertya robusta (Maxim.) Beauverd. Macroclinidium clinidium kpribanum Nakai; P. triloba var. \oribana (Nakai) 



robustum Maxim.; P. macroclinidium Makino Kashiwaba- Makino Sendai-haguma. Median leaves petiolate, alter- 



haguma. Rhizomes creeping, knotty; stems erect, simple, nate, elliptic, about 12 cm. long, about 6 cm. wide, acute, 



30-70 cm. long, thinly puberulent at top; leaves mosdy near cuneate at base, loosely mucronate-toothed, the petioles 2-2.5 



the median portion of the stem, ovate to ovate-oblong, 10-20 cm. long; heads 1-flowered, in panicles. Honshu (near 



cm. long, 7-12 cm. wide, acute, rounded to cuneate at base, Sendai). Hybrid of P. triloba X P. robusta. 



dentate or irregularly incised, puberulent on both sides, ciliate, 



11. AINSLIAEA DC. Momiji-haguma Zoku 



Perennial herbs; leaves petiolate, radical or on the median part of stems, entire to lobed; heads in racemes, spikes, or 

 panicles, often nodding; involucres narrowly tubular; bracts imbricate; receptacles glabrous; florets few, bisexual, fertile or 

 rarely sterile, rarely cleistogamous, the corolla lobes 5, linear, reflexed, the anthers sagittate, the tails connate, usually acuminate; 



