28 BOTANICAL GAZETTE [JULY 
shorter and more truncate, while in the anaphase no cell-plate 
whatever was produced. 
The way in which the chromosomes divide could not be posi- 
tively determined. The segments on entering the nuclear plate, 7 
and even while in the plate as seen from the poles, are much 
curved (fig. 6). A side view of the nuclear plate shows a dense 
mass with projecting arms similar to the corresponding figure in 
the pollen (fig. 6). When the segments move to the poles they 
are seen to be quite long and straight, and never V-shaped as in 
the heterotypic division (fig. 7). After reaching the pole they 
at length fuse, and a membrane is formed around them, thus 
bringing about a truly resting condition again. 
It seems scarcely to be doubted that the process here is 
identical with the second division in the pollen-mother cell. 
Indeed, every appearance in the one has its almost exact coun- 
terpart in the other. 
GROWTH AND DEVELOPMENT OF THE EMBRYO-SAC. 
The further history of the embryo-sac. is very interesting, — 
since it shows some deviations from the ordinary process in 
both monocotyledons and dicotyledons. The two superimposed a 
daughter cells should probably be considered as constituting the : 
so-called ‘axial row” in this case (figs. 6-8). At least, so far : 
as the nuclei are concerned, they are the equivalents of the two : 
cells first formed in Canna. There is therefore a two-celled 
axial row instead of a four-celled one, as in Canna and many — 
other plants. It would be expected then that during develop- — 
ment the lower cell alone would become the embryo-sac, while | 
the upper would undergo dissolution as in nearly all other cases. 
This, however, is not the case. 
At the time when the spindles of the second division occuf, 
the archesporium as described above is two-celled. No walls 4 
are produced by the second spindles, as a result of which each . 
cell now contains two nuclei. This stage is followed by a com : 
paratively long period of growth in which both cells increas€ — 
several times in size, as do also their nuclei, A large number of 
