158 BOTANICAL GAZETTE | SEPTEMBER 
mother cell frequently shows the same color. To be sure some 
preparations show the orange color in connection with the cell 
plate, but the violet of the cell plate itself could nearly always 
be distinguished. The orange color is probably due to the 
small amount of carbohydrate material that sometimes appeats 
at this stage. I do not think that the presence of the above 
mentioned irregular rows of granular trophoplasm is to be take 
as evidence that the spindle fibers break up into granular cyt 
plasm. My preparations show clearly that the spindle fibers a 
entirely distinct from these rows. The history of the fibers 
seems to consist of a gradual shortening and thickening until 
their whole substance is transformed into a cell plate, #.é@ 
membrane. While the above description will apply to thos 
fibers which take part in the formation of the cell plate, the fate 
of those radiating fibers which have no part in this process 
remains unaccounted for. My observations upon this latte 
point have been too limited to base any conclusions upon them. 
In a very early stage in the formation of the cell plate these 
fibers often seem to lose their characteristic radial arrangemet ie 
and to become more of a tangled mass in the cytoplasm (fs: : 
3, 4). It is possible that they become separated from “ 
daughter nuclei and are finally absorbed into the rest if ‘ 
protoplasm. There is one other interesting case to be 20" 
In some ceils of the larch and some of the cells of the root 
of Fritillaria, after the cell plate was formed, a few clearly 
fibers were observed in the region between each daugi 
nucleus and the cell plate. These fibers are probably the ee 
that have been mentioned previously as not showing the ge 
Preparatory to the formation of the cell plate. Their wee 
not determined. Fig. zo is interesting in this conse 
represents a late stage in the division of the pollen mother © : 
when the permanent cell plates are complete, but ease 
still exist numerous fibers all around each nucleus, ome 
into the cytoplasm and reaching in many cases to thet : 
membrane, Whether these are a part of the connecting * 
radiating fibers that existed during the earlier stage 
