372 BOTANICAL GAZETTE | DECEMBER 
beginning. The poles are not diametrically opposite each other, 
but both lie on one side of the nucleus, as shown in fig. 75". Invari- 
ably, too, the axes of the second spindle are at right angles to 
that of the first; that is, though they may be at any angle 
with reference to each other, they always lie in planes which are 
parallel to the first cell plate (figs. 75-18). In the preparations 
selected for drawing, the second spindles are either parallel 
(figs. 14-17, 21), or at right angles (fig. 78). It must not be 
understood, however, that these cases are typical in this respect ; 
they were chosen because in them both the spindles are in view 
at once. 
Undoubtedly the material used for the building up of the 
second spindle is obtained from the disintegration of the first, 
not as fibers, however, but as granules; and in this abundance 
of granular matter ready formed in the cell, we may perhaps see 
an explanation of the rapidity of the second division. 
he phenomena attending the metaphase of the second 
divisions agree completely with those of the first division. 
There is the same absence of mantle fibers (figs. 17; 18), the 
Same continuity of the fibers from pole to pole (fg. 1g), the 
same excess of fibers over the number of chromosomes (/igs. 
18,19). In the anaphase there is the same elongation of the 
chromosomes into crooked or lobulated rods, the same beaded 
appearance of the fibers as soon as the chromosomes have passed 
to the poles (fig 2r), and the same development of secondary 
fibers which run from the poles towards the equator. = 
The beginning of the secondary fibers is shown in fig. ae 
they are more abundant and longer in figs. 20 and 2, In rt 
it is seen that those put out from the poles of one dee ee 
those from the other. The secondary fibers so meeting unite PY 
their ends into continuous threads, which connect the Sas 
of the primary spindles. In this way four secondary SP is : 
are formed; and thus the four daughter nuclei are now aca 
by six symmetrically placed spindles, of which two ae ers 
and four secondary. Meantime, there has been a slight gi ‘is 
tion of the primary spindles so that their planes, which were 
