1905] MOORE—SPOROGENESIS IN PALLAVICINIA 89 
At the completion of the achromatic spindle, the chromosomes 
are found grouped in a ring at the equatorial region of the structure. 
Figs. § and 9 show the arrangement of the chromosomes at this stage. 
Fig. 15 gives a side view, slightly oblique, of the chromosomes at 
metaphase of mitosis. Five chromosomes are in view and the other 
three are hidden or have been removed by the razor in making the 
section. 
I have not been able to make out satisfactorily the details of the 
separation of the daughter chromosomes. The distribution is effected 
very quickly, for great numbers of nuclei in metaphase have been 
observed and a great many in telophase, but very few in anaphase. 
Little indication is given as to the exact manner in which the separa- 
tion takes place. A few instances of chromosomes as they are pulled 
apart are shown in fig. 17. The appearance of the chromosomes 
indicates beyond doubt that they are plastic bodies subjected toa pull, 
and that they are being halved; but what real relation this distribution 
bears to the original tetrads is left in doubt. In fig. 18 we have shown 
anaphase in which the chromosomes are somewhat scattered upon 
a very broad spindle. There are five near each pole and one almost 
half way between. It is evident that the remaining chromosomes are 
upon another section. 
During telophase the chromosomes are found arranged in compact 
rings at the two poles. When one end of a spindle abuts on a dividing 
wall between two lobes, the ring at that end sometimes lies very close 
to this wall, partially surrounding it (fig. 19). 
There is no resting stage between the first and second mitoses. 
The chromatic elements of the nucleus do not resolve themselves 
into a reticulum and the chromosomes do not lose their individuality. 
The rings of chromosomes which have been formed at the telophase of 
the first division merely alter their positions, so that their planes 
lie at right angles to one another. It is evident from jig. 20 that the 
chromosomes come in contact and form a thick spirem, but do not 
lose their identity. This is the nearest approach to a resting stage I 
have been able to find, and I believe it is unusual for the reconstruction 
of the nucleus to proceed even this far. No nuclear membrane is 
formed at the end of the first mitosis and no cell plate is laid down. 
In a few instances granules were seen across the equatorial portion 
