456 
flattened dorsiventrally, and possess many long rhizoids. 
The reproductive organs are developed on the flattened 
monoecious. 
BOTANICAL GAZETTE 
[DECEMBER 
They are 
dorsal side and in their organization differ essentially from those of Botry- 
chium virginianum as described by JEFFREY.3 
The tig ila upon segmentation does not develop directly into a 
1G. 1.—Photomicrograph of a section through 
- ee and young sporophyte of Sceptri- 
“um £6 The section is vertical, and trans- 
verse Fie gametophyte. The root is already 
Senses from the under side of the gametophyte, 
while! the position of the first leaf was marked by a 
pronounced elevation on the upper side. a, arche- 
gonium; s, suspensor; ¢, stem tip; /, first leaf; r, 
Toot. X_60. 
spherical protocorm, as is 
the case in all other ferns 
which have been studied, 
but first gives rise to a long 
suspensor, which burrows 
into the tissue of the game- 
tophyte in the manner 
characteristic of certain 
lycopod embryos. At the 
tip of this suspensor a 
spherical protocorm is or- 
ganized, out of which the 
stem and root apices are 
shortly differentiated. The 
axis of the metacorm trans- 
fixes the protocorm and all 
the tissue of the latter, ex- 
cept the suspensor, becomes 
a permanent part of the 
metacorm. The embryo 
does not, therefore, possess 
a lateral cotyledon (nurs- 
ing-foot) as does Botry- 
chium virginianum. The 
root grows downward and 
emerges jrom the under side 
of the gametophyte, and at 
a later period the first leaf 
breaks through the upper 
surface. The relation of 
the members in the young 
embryo and its orientation in the gametophyte are well illustrated by the 
accompanying figure. 
A study of the mature sporophytes of the ternate species of Botrychium 
3 Univ. of Toronto Studies 1:1-32. 1898. 
