VOLUME XXIX NUMBER 2 
BOTANIGAL:G4AZE¢7TE 
FEBRUARY 1900 
STUDIES IN THE DEVELOPMENT OF THE POLLEN 
GRAIN IN SYMPLOCARPUS FCETIDUS AND PEL- 
TANDRA UNDULATA. 
B. M. Duccar. 
(WITH PLATES I AND II) 
Deraits of the nuclear and cell division in the development 
of the pollen grain in these plants were first studied in Symplo- 
carpus fetidus, an abundance of favorable material having been 
secured and satisfactorily preserved during the winter of 1897-8. 
When it afterwards seemed well to study another plant in the 
same order, Professor Atkinson kindly put at my disposal mate- 
tial of Peltandra undulata which he had collected two years ear- 
lier. The latter material was also well fixed and preserved ; but 
since it gave fewer developmental phases, Peltandra has been 
used largely by way of comparison with Symplocarpus. In gen- 
eral, these two plants agree in important phases of the cell activ- 
ities, but there is considerable variation in some details. 
The origin of the primitive archesporium was not studied in 
either plant, but in Peltandra there were found stages showing 
the divisions in the archesporial cells. These divisions are of 
the normal vegetative type; the number of chromosomes present, 
moreover, is that characteristic of the sporophyte. 
THE RESTING ARCHESPORIAL CELL AND ITS NUCLEUS. 
In Symplocarpus the cells of the definitive archesporium pass 
the late fall and early winter months in the resting condition. 
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