116 BOTANICAL GAZETTE [FEBRUARY 
(fig. 65). The early divisions of the endosperm nuclei occur 
more rapidly than those in the embryo, for while there are yet 
but two cells in the embryo, the endosperm nuclei have passed 
to their third division, a condition shown in jig. 66. The 
antipodal cells are still to be seen in a small pocket at the 
extremity of the sac. Their final disappearance does not occur 
until a considerably later stage. 
It is customary to speak of the cell which produces the 
octants as the embryo cell, and of all the remaining cells as 
belonging to the suspensor. The term is never strictly accurate, 
for even in such cases as Capsella the dermatogen and periblem 
of the root are contributed by the cell below the octants. Again 
some authors, noticing that the first wall in this ‘embryo cell” 
is usually longitudinal, conclude that, conversely, the first longi- 
tudinal wall identifies the embryo cell —a definition which breaks 
down in cases where the octants are formed by a transverse wall 
followed by two longitudinal walls, as has been claimed by 
Schwere (24) to be the rule for Compositae. If, however, we 
give to the term ‘embryo cell”’ a uniform signification, namely, 
the cell which produces the octants, we shall at least justify its 
use as a term of convenience. 
Another term of varied application is the word hypophy sis, 
or “‘Anschlusszelle.” In Hanstein’s account of the embryo of 
Capsella (2), the cell next below the octants divides by 4 
transverse wall, and its upper half divides again to form the 
‘‘Periblem-schlusszelle” and the ‘‘ Dermatogen-schlusszelle,’ * 
the latter, by repeated splitting, forming the layers of the root 
cap. Hanstein applies the term hypophysis sometimes t0 
this original suspensor cell, at other times more particularly to 
its upper half, thus designating as the ‘“Anschlusszelle” the cell 
which actually completes the embryo. On the other hand 
Schwere, for example, applies the term to the cell which is, 
next below the embryo cell, indeed, but which only partially 
*Throughout the rest of this account the rather cumbrous words “ Dermatoge®” 
schlusszelle” and “ Periblem-schlusszelle” will be translated by the terms dermatogen 
and feriblem terminals, respectively. The fact that they later become the dermatoge? 
and periblem initia/s need cause no confusion when the latter terms are substituted. 
