304 BOTANICAL GAZETTE [May 
finally dissolves, leaving the two groups of daughter chromo- 
somes quite apart (fig. 72), after which each daughter nucleus 
proceeds to organize its new membrane. The intervening space, 
at first vacuolate, becomes filled with granular cytoplasm and 
the mitosis is finished. 
There appears to be no positive evidence that the mitoses 
described above for the oogonium and antheridium are reducing 
divisions, whatever may be the speculations upon that point. 
Little can be said on this subject in our present very incomplete 
knowledge of the mitotic figures at other periods in the life 
history of Albugo. The study of the oospore may lead to some 
important results, although the writer’s incomplete observations 
were no more promising than the examinations of Wager (96), 
Berlese (98), and Stevens (99). An investigation of this sub- 
ject is much to be desired, and should include the study of the 
nuclear figure at some period of vegetative activity. 4. candida, 
however, is not the most favorable subject for such an examina- 
tion, as the nuclei are small. A. portulacae or A. bliti would be 
more satisfactory. 
THEORETICAL CONSIDERATIONS. 
The writer must confess a feeling of considerable uncertainty 
as to the cytological conditions and perhaps homologies of the 
sexual organs found in certain Phycomycetes. Much interesting 
material is likely to be presented through future investigations 
in this field, and particularly upon those forms whose gametes 
are well known to be multinucleate. Accordingly any review of 
the present situation can only offer suggestions of what may 
later become established by research. The subject is a very 
interesting one, but the difficulties of investigation are unusual, 
not only on account of the necessary technique, but in the nature 
of the material that must be studied. 
It is well known that the gametes of certain fungi contained 
in the Mucorales are multinucleate. Several investigators have 
studied these structures, and the zygospore which results frou: 
their fusion, but none have given a clear account of the cytologr 
