316 BOTANICAL GAZETTE | MAY 
sporophore which originated from a single erect hypha. The 
two lower cells have sent out branches, which, together with the 
original hypha, have formed a stalk bearing a small head of 
gonidia at its summit. The young branches are hyaline at the 
tips, but brown in the older portions. They soon become merged 
with the other hyphae forming the stipe. 
In the more complex forms of the small gonidiophores there is 
seen an approach to the perfect sporophores of Trichurus. In 
fact, they are identical with the latter both in structure and 
manner of formation, differing only in the number of hyphae 
which go to make up the stipe arid head. It is easy to forma 
complete series, showing all gradations from the simplest hypha 
bearing a single chain of gonidia to the most complex sporo- . 
phore with several hundred spore-chains. 
At a time varying from four to six days after sowing, the 
vegetative hyphae have formed a dense mat of mycelium in the 
medium in which they are growing. At the center of the colo- 
nies the threads become so numerous and interwoven that they 
form almost a stroma. From this stroma-like mat the perfect 
sporophores arise, either singly or in small clusters. They first 
appear as stout fascicles of hyphae, which arise perpendicular to 
the substratum. The growing points of the threads are hyaline, 
but in the older parts of the bundle they assume the characteris- 
tic brown color of thestipe (fig. 78, of Stysanus stemonites).4 The 
threads branch in the manner described above, the branches 
remaining closely appressed to the bundle and growing with it. 
During longitudinal growth the central threads are in advance of 
the peripheral hyphae, giving the bundle a linear, pointed form. 
Before the bundle has completed its growth, the individual 
hyphae begin to send out small, curved branches or basidia at 
some distance from their growing points. The basidia are 
thus formed in centripetal order. Those first formed begin to 
bear chains of spores before the bundle or stipe has completed 
4The formation of the stipe and head in the two forms, 7richurus — yea 
Stysanus stemonites, is exactly alike, and it was not considered necessary to Serr wet 
this stage more than once. Figs. 78 and sg are from Stysanus, but they will ans 
equally well for Trichurus. 
