320 BOTANICAL GAZETTE [MAY 
to change the external appearance of the colony (jigs. 75, 
16,17). The gonidiophores themselves, on the other hand, are 
very much alike in the two plants, as figs. 75, 76 and 17 will 
show. In Stysanus also we can find a complete series showing 
all gradations between these small gonidiophores and the perfect 
sporophores. 
The mature sporophores are small, rarely attaining a height 
of 0.75™. They arise either singly from the center of the col- 
ony, or several arise together from a stroma-like base. The 
stipe elongates rapidly, and soon begins to bear basidia and 
chains of gonidia which remain together in a tangled head so 
long as they are dry, but when placed in water they break apart 
instantly. Such sporophores from which most of the gonidia 
have fallen away are shown in figs. 20 and 27. 
CONCLUSION. 
From this study there can be no doubt of the close relation- 
ship of Trichurus spiralis and Stysanus stemonites. Not only are the 
perfect forms similar to each other, but they agree closely in 
their entire development, which may be summed up as follows. 
The spores germinate in a peculiar manner by means of a stout 
primary germ tube or proembryo. The mycelium forms a small, 
compact colony, from which there arises a gonidial fructification 
very different from the normal fructification of the plants. The 
normal fruit consists of a bundle of hyphae bearing basidia and 
chains of gonidia at the summit. All gradations between the 
simple sporophores and the compound heads exist. The char- 
acters which distingish these two plants from each other are the 
presence of tortuous sterile threads in the head of Trichurus and 
the difference in the size of the plants and spores. Among 
these low forms these characters are, perhaps, sufficient ive 
generic separation. The plant treated in the first part of this 
paper has provisionally been referred to the genus Trichurus 
Clements & Shear. Trichurus was characterized by them thus: - 
‘‘As in Stysanus, but the capitulum densely beset with long 
strict bristles.” The present plant, however, differs from this 
