130 BULLETIN : MUSEUM OF COMPARATIVE ZOOLOGY. 



It is not clear, then, whether the galea develops from the outer or 

 the inner lobe of a bii'amous appendage, although Uzel's account is, at 

 least, not inconsistent with his description of the first maxilla, which I 

 have already criticised. Although Uzel does not state as much, his 

 figures indicate the palpus to be an appendage of the primary funda- 

 ment, as it is in Anurida. In this genus, however, no third branch ap- 

 pears, as I have said ; but, from analogy with the fii'st maxilla, the inner 

 of the two branches represents undiflerentiated galea and lacinia. 



The rotation in a frontal plane of the second maxillary fundament of 

 Campodea — which does not occur in Anurida — enables me to homo- 

 logize the finished labium of Campodea with the apparently different 

 labium of all other insects. If Uzel's figures are compared with Figure 

 12 of V. Stummer-Traunfels ('91), it is easy to see that the embryonic 

 structures by Uzel designated limx^ (lacinia), lemx^ (galea) and j^i^^i 

 (palpus) are with hardly a doubt respectively represented in the adult by 

 the parts which v. Stummer-Traunfels termed iqy. (" untere INIundplatte "), 

 pi. (" tasterformige Papille ") and pj^- (" Tastwarze "). These homo- 

 logies, however, could never have been settled upon merely anatomical 

 grounds. 



What Grassi ('86^, Tav. IV. Figura 3), then, considered to be the 

 under lip {la. in.) of Campodea is but the anterior part of the true 

 labium; the "labial palpi" (pa. li.) are really gale?e borne upon a 

 region representing the mentum, and the " labial papillae " {pa. la.) 

 are but modified palpi. As in Collembola, the labium is anteriorly and 

 deeply cleft. 



Japyx is so close to Campodea that the same conclusions may doubt- 

 less be applied to both genera. In Japyx the labium, as in Collembola, 

 is split and bears a median sulcus (Grassi, 86^ Tav. III. Figura 21) 

 much like that of Orchesella (Folsom, '99, Plate 4, Figure 29). Ex- 

 amining Figure 1 of v. Stummer-Traunfels ('91, Taf. II.), the lacinia and 

 galea are clearly represented, as in Campodea ; the true palpus, however, is 

 but obscurely differentiated in the region behind the so-called palpus (pi.) 

 and nearer the median plane. The eversible papilla; of the anterior part 

 of the labium, as described by Meinert ('67, p. 3G9) and Grassi ('86'', 

 p. 31), are probably honiologous with the papilla3 of Orchesella which I 

 designated pip. ('99, Plate 3, Figure 24). 



For Lepisma, Heymons (97% p. 590, Figur 11) gives, first, a pair of 

 simple second maxillary fundaments and later (Taf. XXX., Figur 20) a 

 long palpus with a small, basal, inner lobe, and states (p. 592) " Die 

 Lobi oder spatei-en Ladentheile der Maxillen sind in diesen Stadien erst 



