204 BULLETIN : MUSEUM OF COMPARATIVE ZOOLOGY. 
which had formed only one polar cell, the cleavage cells contained 84 
double chromosomes. 
As the eggs of the second type were rare and sometimes showed multi- 
polar spindles, Brauer is uncertain whether they were really capable of 
normal development or not. Petrunkewitsch is certain that they must 
have been purely pathological, for he never observed evidence of any 
such second method of maturation in his own preparations, though this 
was the especial object of his search, and he worked with material from 
the same locality, Triest, that had furnished Brauer’s material, and in 
addition with material from a second locality, Odessa, where male Arte- 
mias not infrequently occur. 
But a moment’s reflection will show that the apparently discordant 
results of Brauer and Petrunkewitsch are readily reconcilable. Brauer’s 
second type of maturation may have been observed in the rare male (or 
fecundable) eggs. 
But why, then, it may be asked, did not Petrunkewitsch encounter 
this second type of egg, the especial object of his search, for he exam- 
ined material from Odessa, where males frequently occur. Probably be- 
cause he, as he explicitly states, worked erelusively with winter eggs 
(‘‘ Dauereier ”), whereas Brauer worked both with summer eggs (“ Subi- 
taneier”’) and with winter eggs. Though Brauer makes no statement 
concerning the matter, I confidently hazard the conjecture that the 
second type of maturation was observed by him only among the summer 
eggs, for in no species, so far as I know, in which parthenogenesis 
occurs, has the development of a male animal from a winter egg ever 
been observed. In parthenogenetic Crustacea, Rotifera, and Platodes 
alike, there invariably hatches from the winter egg a parthenogenetic 
female. Should Petrunkewitsch study the parthenogenetic summer eggs, 
instead of the winter eggs, produced by Artemias of the Odessa race, I 
venture to predict that his search for the second type of maturation 
will be abundantly rewarded, at least to this extent, that he will 
find the occurrence of two maturation divisions in the male summer 
eggs. 
It is doubtful whether the other process observed by Brauer, a fusion 
of the nucleus of the second polar cell with the egg nucleus, takes place 
in the development of the male Artemia. More probably the result of 
this process would be the same as that of fertilization, or of an entire 
suppression of the second maturation division; namely, the production 
of a female in which the male character is recessive. This view is 
quite in harmony with Brauer’s own interpretation of his observations. 
