360 BULLETIN : MUSEUM OF COMPARATIVE ZOOLOGY. 
has found what he calls “ Kornchenkugeln.” Judging from his figures 
of them, they do not look much like the “ Kornchenkugeln” of the 
Muscidae, nor does their migration into the lumen of the intestine agree 
with what has been found in Diptera. Moreover, he states that these 
phagocytes are not numerous enough in the region of the midintestine 
to account for the degeneration of the muscles of this region, and conse- 
quently infers that there is chemical degeneration as well as phagocyto- 
sis. Such different methods of degeneration in similar muscles of the 
same animal is improbable. But the principal reason for believing that 
there is no phagocytosis of these muscles in Thymalus and other Cole- 
optera lies in the exact similarity of all their changes to those occurring 
in the muscles of the leg type. In these muscles it can be stated with 
certainty, not only that there is no phagocytosis, but also that the 
larval muscles metamorphose into the imaginal muscles instead of 
degenerating. 
The typical “ Kornchenkugeln” which Deegener finds, but which 
Rengel could not find, are met with in Thymalus. That is to say, there 
are to be found leucocytes containing bodies many of which would 
answer the description given by Deegener, but these leucocytes are not 
such “ Kérnchenkugeln” as Weismann: found, This is evident from 
some of the appearances reproduced in Figures 40-48 (Plate 7). 
These all represent leucocytes found in old pupae magnified 1600 
diameters. Figures 43 and 46 look like leucocytes containing de- 
generating nuclei, and there is a possibility that such may be the true 
explanation of some of them; none of them, however, are nuclei from 
the intestinal muscles. Figures 40, 42, and 47 show inclusions which 
certainly are not degenerating nuclei, and since there are found transi- 
tional stages (Figure 48) to the first mentioned conditions, it is probable 
that all of the inclusions are of the same kind. The most probable 
interpretation of them is that they are intracellular parasites. This 
view is strengthened by the presence of apparently similar bodies in 
the intestinal epithelium of resting larvae. Also, bodies similar to the 
deeply stained portions of Figure 40 are found very numerously in the 
body cavity and lumen of the intestine of old pupae and young imagines. 
The true nature and relationship of these bodies cannot be stated with 
certainty as yet, but whatever they may be, very few, if any of them, 
can be called “ Kornchenkugeln.” 
Concerning the formation of the intestinal muscles of the imago, my 
observations, again, are in harmony with those of Rengel, and disagree 
with those of Deegener. The reconstruction of the intestinal muscles 
