RESULTS FROM THE MAMMAL SURVEY. 405 



they agree on an exaggerated scale more with those of certain 

 species of Fjpimys (e.g., E. macleari and hlanfordi), with some 

 approach to those of Lenothrix. 



M^ with the laminee strongly curved and zigzagged (snggesting 

 Lenomys) ; inner cusps on first tM^o laminse large, triangular, witii 

 projecting points behind, their tips sticking up high above the well 

 marked valley between them and the median cusps ; no internal or 

 "x" cusps on the third lamina; external cusps little developed, 

 about as in Ejnmys macleari. M" with antero-internal cusp, large, 

 antero-external obsolete. M^ with large antero-internal cusp, and two 

 equally projecting salient angles on the internal side behind that 

 cusp. Lower teeth withoiit special peculiarities. Root of lower 

 incisors not projecting in a capstile on the outer side of the mandible. 



Genotj^pe. — Dacnomys millardi, sj). n. 



The proper position of this giant rat is somewhat dou.btful. At 

 first I supposed it to be near Lenomys and Lenothrix, the angularity' 

 of the molars and the presence in them of the characteristic connect- 

 ing points between the laminae being somewhat as in those genera, 

 to which however there is no external resemblance. But Mr. 

 Hinton has kindly drawn my attention to the fact that cei'tain 

 Eastern species of LJpimys, notably E. macleari and hlanfordi, have 

 in different degrees, a more or less similar molar structure, Dacnomys 

 differing from these mainly by the disproportionate size of the teeth, 

 and the reduction of the zygomatic plate. It would thus appear 

 to form another link in a chain of forms ranging in molar structure 

 from the simplified Epimys rattus type, through {V) E. macleari and 

 its allies, (2) Dacnomys, and (3) Lenothrix and Di-plothrix to (4) 

 the genera with such highly complex zigzagged teeth as those of 

 Lenomys, Grateromys and Mallomys. Concurrent with the greater 

 complexitj^ of the molars there is a reduction in the anterior projec- 

 tion of the zygomatic plate, and except in the two last named 

 genera, an increase in the heaviness of the supi-aorbital and parietal 

 ridges. As a ready means of diagnosis it ma}^ be noted that in 

 every Ejjimys, the length of the teeth goes more than twice in the 

 distance from the back of M^ to the back of the condjde, while in 

 Dacnomys it goes less than twice. 



Dacnomys millardi, sp. n. 



A large plain looking brown rat with unicolor tail. Size about 

 as in a small bandicoot. Fur of normal character, rather short and 

 thin, hairs of back about 15-16 mm. in length; the longer piles not 

 excessively elongated. General colour above near " olive brown", 

 finely lined with blackish and grizzled with dull buff3^ Sides rather 

 lighter. Belly pale brownish, not sharply defined from the colour 

 of the flanks, the hairs pale slaty brown at base, dull cream}- white 



