160° BOTANICAL GAZETTE [MARCH 
ten cells in the endosperm, but the endosperm does not continue 
to keep pace, and very soon the cells of the embryo outnumber 
the nuclei of the endosperm. The nuclei of the endosperm in 
Salix are never separated by cell walls. 
DEVELOPMENT OF THE EMBRYO. 
The first division of the oospore is always transverse to the 
longer axis of the embryo sac (figs. gz, 42). Occasionally this 
division separates the oospore into approximately equal parts, but 
it is more usual to find the suspensor cell larger and somewhat 
tapering, while its sister cell, which gives rise to the greater part 
of the embryo, is uniformly hemispherical. The suspensor cell 
divides transversely, and the daughter nuclei pass into the resting 
stage with full sized nucleoli before the embryo cell divides (fig.43)- 
The first division of the embryo cell is always longitudinal 
(jigs. 44-46). The literature of the subject indicates that this 
division is almost universal in angiosperms, if we except those 
which have no suspensor and those in which the suspensor, 
though present, contributes nothing totheembryo. For instance, 
in Capsella after the first division of the oospore, the cell nearer _ 
the micropyle undergoes several divisions, forming the long” : 
suspensor, while its sister cell remains passive until the first 
longitudinal division occurs. This seems to be mere ee 
but it is quite probable that in dicotyls the terminal cell in which — 
the first longitudinal division appears gives rise to the greater 
part of the embryo. Vines in his Zext-Book has unfortunately 
figured the first division of the embryo cell in the type PT 
as transverse. The figures are uauneqensine ‘after Goebel and — 
_ Hanstein” but Hanstein (3) figures the first division as vertical, 
and Goebel has followed him. Vines’ text, however, without any — 
particular reference to Capsella, states that the first division is 
usually longitudinal. Some of Hanstein’s figures, like his figs. 9 
and zz, which show a complete differentiation of the dermatogen — 
before the first vertical division, certainly need. to be ve rift 
especially since the drawings were made from embr c 
oe from the ovule and rendered transpe rer 
