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1897 | LIFE HISTORY OF LILIUM PHILADELPHICUM 425 
It is unfortunate that the terms cyanophilous and erythro- 
philous are becoming established, since the affinity is for basic or 
acid dyes, and not for blue or red colors. That the terms are 
misnomers becomes evident when a combination like safranin 
(basic) and acid green (acid) is used, for the cyanophilous 
structures take the red, and the erythrophilous the green. 
THE MOTHER CELLS AND TETRADS. 
Fairly complete series were obtained in Z. tigrinum and L. 
Philadelphicum, but since the two species showed very similar 
results only ZL. figrinum is described. 
The mother cell develops its tetrads after the usual manner 
among monocotyledons. It was my original purpose to make a 
cytological study of these cells, chiefly with reference to the 
phenomena involved in the ‘“ reduction division,” but my atten- 
tion was diverted to certain structures of the mature spore, 
which will be hereafter described. However, certain cytological 
notes obtained may be of interest. 
The nuclei of the mother cells in early spirem stages show a 
single much twisted ribbon with a row of chromatin granules on 
each edge. In many cases it could be seen that the chromatin 
granules were arranged in opposite pairs (figs, rand ra). These 
pairs are separated by a longer stretch of ribbon than is figured 
by Guignard in his description of L. - Martagon. The granules 
are usually more or less ellipsoidal i in shape, the longer axis 
coinciding with that of the ribbon. With cyanin and erythrosin 
the ribbon stains red, and the granules blue. The ribbon splits 
longitudinally throughout its entire length before it segments 
into chromosomes (fig. 1). The nuclei showed twelve segments 
of this double thread in all cases in which the number was 
definitely ascertained. The further history of the chromosomes 
and the formation of the spindle were not followed. 
In the tetrad stage the nuclear thread is not nearly so intri- 
"cate, and is often spirally wound inside the nucleus, somewhat 
_ilikea chromatophore of Spirogyra (fig. 2.) In many cases it 
seemed as if even in ree Se a the ee of the future 
