456 BOTANICAL GAZETTE [ JUNE: 
some-like bodies or by the changed position of the nucleus the 
writer has not yet been able to surely determine. They have 
meanwhile greatly increased in size, now measuring usually from 
18 to 20 in diameter. They now have a clearly distinguishable 
outer wall of considerable thickness and the contents are evenly 
and beautifully vacuolate. 
When the nuclear spindle is formed these bodies always take 
up a position directly opposite the polar ends. In the monaster 
stage of the division (fig. 7) the spindle, which occupies a trans- 
verse position both to the pollen tube and to the elliptical nucleus, 
is composed of fine kinoplasmic filaments and seems to be entirely 
within the still preserved nuclear wall. It is blunt poled, if indeed 
it is not multipolar, and does not seem to have any kinoplasmic 
connection with the centrosome-like bodies. In the resting con- 
dition of the nucleus the centrosome-like bodies were surrounded 
by radiating threads of kinoplasm, but no indications of these can 
now be seen. There is, however, in most cases a slight radial 
arrangement of the protoplasm immediately bordering the wall. 
The structure of the bodies has meanwhile undergone consider- 
able change (jigs. 7,8). The wall swells up and apparently sep- 
arates into fragments which in cross-section at this stage show as 
abroken line. The vacuolated contents meanwhile contract away 
from the wall, leaving a clear space intervening, which is, how- 
ever, traversed by a few slender filaments. The body at this stage 
presents the appearance of disintegration. 
When the cell-plate is formed (fig. 9) the centrosome-like 
body is found entirely broken up, the fragments appearing in the 
_ cytoplasm as a number of granules mixed with plates or mem~ 
_ branes which appear to be fragments of the wall. After the new 
ae cell wall is completed and the daughter nuclei have returned 
_ to the resting stage, showing a nucleolus, the fragments of the 
broken-down — centrosome-like body present the appearance 
i. : shown i in fig.t sib A little cluster of Rae appear in the exact 
o : a location ¢ of t t , which stain deeply and often 
a ue “ie be two in | number, have separated and pared toward 
