18 BOTANICAL GAZETTE [JANUARY _ 
determine. It seems highly probable, however, that, like the 
internodal cells of Chara and Tradescantia, they may have lost 
the capacity to divide, the increase in number being entirely 
provided for by the division of the binucleate cells. The devel- 
opment of the carpophore, as pointed out by De Bary (4, p. 
49) and others, is no doubt to be interpreted as consisting in the 
which are to be traced back through the pileus into the stipe, 
and so connected with the mycelial branch or branches from 
which the carpophore sprung. The growth would then be 
largely apical for these hyphae, the older cells lying in the 
pileus and stipe, and the youngest, latest. formed, in the hymen- 
ium. That all the hyphae of the stipe have end cells in the 
hymenium is, of course, not necessarily assumed, though it is not 
improbable, in view of the wide expanse of the latter, that this 
is the case. 
What the condition of the mycelial cells is in the species of 
Coprinus studied I have not been able to determine. Maire 
says that the mycelium of Coprinus radiatus is made up of uni- 
nucleated cells. The base of the stipe of C. ephemerus is sut- 
rounded by a mass of hyphae with multinucleated cells ; whether 
these, however, represent the condition of the primitive mycel- 
ium from which the carpophore sprung I am by no means sure. 
It may well be that in the work of collecting and transporting 
dition represented in the basidiospore may be perpetuated for 
longer or shorter periods of development in different cases. 
